Dunes near The Hague, February
Ammophila (synonymous with Psamma P. Beauv.)is a genus consisting of two or three very similar species of grasses; common names for these grasses include Marram Grass, Bent Grass, and Beachgrass. These grasses are found almost exclusively on the first line of coastal sand dunes; their extensive systems of creeping underground stems or rhizomes allow them to thrive under conditions of shifting sands and high winds. Ammophila species are native to the coasts of the North Atlantic Ocean where they are usually the dominant species on sand dunes. Their native range includes few inland regions, with the Great Lakes of North America being the main exception. The genus name "Ammophila (Am-mó-phi-la) " originates from the Greek words of Ammos (ἄμμος), meaning sand, and Phillia (ϕιλος), meaning lover.
The Ammophila grasses are widely known as examples of xerophytes, which are plants that can withstand arid conditions such as deserts or sandy beaches. Its xerophytic adaptations (mentioned below) allow it to thrive under conditions most plants could not survive. Despite their occurrence on seacoasts, Ammophila grasses are not particularly tolerant of saline soils; they can tolerate a salinity of about 15 g/l (1.5%), which makes them "moderate halophytes".
Ammophila builds coastal sand dunes and thus stabilizes the sand. For this reason, the plants have been introduced far from their native range. Alfred Wiedemann writes that, starting in the early 1800s, Ammophila arenaria "has been introduced into virtually every British colonial settlement within its latitudinal tolerance range, including southeast and southwest Australia, New Zealand, South Africa, the Falkland Islands, and Norfolk Island. It has been planted widely in Japan and has been reported from Argentina and Chile." Ammophila species were introduced in the late 1800s on the Pacific coast of North America as well, and massive, intentional plantings were continued at least through 1960. In essentially all of the locations where they have been introduced, Ammophila plants are now listed as invasive, and costly efforts are underway to eradicate them.
Only two species seem incontrovertible: A. arenaria and A. breviligulata. Two other species have been proposed, and are discussed below.
In Europe, Ammophila arenaria has a coastal distribution, and is the dominant species on sand dunes where it is responsible for stabilising and building the foredune by capturing blown sand and binding it together with the warp and weft of its tough, fibrous rhizome system. Marram grass is strongly associated with two coastal plant community types in the British National Vegetation Classification. In community SD6 (Mobile dune) Ammophila is the dominant species. In the semi-fixed dunes (community SD7), where the quantity of blown sand is declining Ammophila becomes less competitive, and other species, notably Festuca rubra (Red Fescue) become prominent.
The ability of marram grass to grow on and bind sand makes it a useful plant in the stabilization of coastal dunes and artificial defences on sandy coasts. The usefulness was recognized in the late 18th century. On the North Sea coast of Jutland, Denmark, marram grass was traditionally much used for fuel, thatch, cattle fodder (after frost) etc. The use led to sand drift and loss of arable land. Hence, legislation promoting dune stabilization came into force in 1779 og 1792, successively leading to system of state-supported dune planters overlooked by dune bailifs. Marram grass was - and still is - propagated by root and shoot cuttings dug up locally and planted into the naked sand in periods of relatively calm and miost weather.
Marram grass has been widely used for thatch in many areas of the British isles close to the sea. The harvesting of marram grass for thatch was so widespread during the 17th century that it had the effect of destabilizing dunes, resulting in the burial of many villages, estates and farms. In 1695 the practice was banned by an Act of the Scottish Parliament:
Considering that many lands, meadowes and pasturages lying on sea coasts have been ruined and overspread in many places in this kingdom by sand driven from adjacent sand hills .... His Majesty does strictly prohibit and discharge the pulling of bent, broom or juniper off the sand hills for hereafter.
European Marram Grass (left) with Velvet Bent (right).
European Marram Grass at Oostduinkerke, Belgium.
American beachgrass on Lake Michigan in November.
Classification System: APG II (down to family level)
Species: A. arenaria - A. arundinacea - A. baltica - A. breviligulata - A. brevipilis - A. champlainensis - A. curtissii - A. littoralis - A. longifolia - A. pallida - A. villosa
|For more multimedia, look at Ammophila on Wikimedia Commons.|