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His research has focused on the use of dung-feeding beetles (Scarabaeidae: Aphodiinae, Scarabaeinae; Geotrupidae; Hybosoridae) as indicators of environmental disturbance.
His work was centered on tropical forests in Malaysia, examining the community dynamics of dung-feeding beetles in primary and derived forest ecosystems.
His particular interests include canopy communities where a new guild of arboreal dung beetles was found during work in northern Borneo, revealing a dung-rolling species of Onthophagus.