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Domestic cat[1]
Conservation status
Scientific classification
Kingdom: Animalia
Subkingdom: Eumetazoa
Phylum: Chordata
Subphylum: Vertebrata
Class: Mammalia
Subclass: Theria
Order: Carnivora
Suborder: Feliformia
Family: Felidae
Subfamily: Felinae
Genus: Felis
Species: F. silvestris catus
Binomial name
Felis catus
(Linnaeus, 1758)[2]
Felis catus domestica (invalid junior synonym)[3]
Felis silvestris catus[4]
The cat (Felis silvestris catus), also known as the domestic cat or housecat[5] to distinguish it from other felines and felids, is a small domesticated carnivorous mammal that is valued by humans for its companionship and its ability to hunt vermin and household pests. Cats have been associated with humans for at least 9,500 years,[6] and are currently the most popular pet in the world.[7] Due to their close association with humans, cats are now found almost everywhere on Earth. This extreme adaptability and their worrying impacts on native animals has led to them being classed as an invasive species. Most of these problems are caused by the large number of feral cats worldwide, with a population of up to 60 million of these animals in the United States alone.
Cats are similar in size and anatomy to the other Felids, with light, flexible bodies and teeth adapted to killing small prey. A skilled predator, the cat hunts over 1,000 species for food, using its excellent eyesight and hearing. Unusually, cats have lost the ability to taste sugar and in some breeds show hereditary deafness. Despite being solitary hunters, cats are a social species and use a variety of vocalizations, pheromones and types of body language for communication. These include meowing, purring, trilling, hissing, growling, and grunting.[8] They are also bred and shown as registered pedigree pets. This hobby is known as cat fancy.
As The New York Times wrote in 2007, "Until recently the cat was commonly believed to have been domesticated in ancient Egypt, where it was a cult animal."[9] A study that year found that the lines of descent of all house cats probably run through as few as five self-domesticating African Wildcats (Felis silvestris lybica) circa 8000 BC, in the Near East.[4] The earliest direct evidence of cat domestication is a kitten that was buried with its owner 9,500 years ago in Cyprus.[10]


Nomenclature and etymology

The word cat derives from Old English catt, which belongs to a group of related words in European languages, including Welsh cath, Spanish gato, Basque katu, Byzantine Greek Κάτια kátia, Old Irish cat, Frisian and Dutch kat, German Katze, and Old Church Slavonic kotka. The ultimate source of all these terms is Late Latin catus, cattus, catta "domestic cat", as opposed to feles "European wildcat". It is unclear whether the Greek or the Latin came first, but they were undoubtedly borrowed from an Afro-Asiatic language akin to Nubian kadís and Berber kaddîska, both meaning "wildcat".[11] This term was either cognate with or borrowed from Late Egyptian čaus "jungle cat, African wildcat" (later giving Coptic šau "tomcat"),[12] itself from earlier Egyptian tešau "female cat"[13] (vs. miew "tomcat").[14] The term puss (as in pussycat) may come from Dutch poes or from Low German Puuskatte, dialectal Swedish kattepus, or Norwegian pus, pusekatt, all of which primarily denote a woman and, by extension, a female cat.[15] This name could be related to Hindi कथि or Turkish qadi. [16]
A group of cats is referred to as a "clowder", a male cat is called a "tom" (or a "gib", if neutered), and a female is called a "molly" or "queen". The male progenitor of a cat, especially a pedigreed cat, is its "sire", and its female progenitor is its "dam". An immature cat is called a "kitten" (which is also an alternative name for young rats, rabbits, hedgehogs, beavers, squirrels and skunks). In medieval Britain, the word kitten was interchangeable with the word catling. A cat whose ancestry is formally registered is called a pedigreed cat, purebred cat, or a show cat. In strict terms, a purebred cat is one whose ancestry contains only individuals of the same breed. A pedigreed cat is one whose ancestry is recorded, but may have ancestors of different breeds. Cats of unrecorded mixed ancestry are referred to as domestic longhairs and domestic shorthairs or commonly as random-bred, moggies, mongrels, or mutt-cats.

Taxonomy and evolution

The wildcat Felis silvestris is a close relative of the domestic cat.
.The Felids are a rapidly evolving family of mammals that share a common ancestor only 10-15 million years ago,[17] and include in addition to the domestic cat, lions, tigers, cougars, and many others.^ By on November 15, 2009 at 10:17 pm chinese food recepies… .
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Within this family, domestic cats (Felis catus) are part of the genus Felis, which is a group of small cats containing seven species.[1][18] Members of the genus are found worldwide and include the Jungle Cat (Felis chaus) of southeast Asia, the African Wildcat (Felis silvestris lybica), the Chinese Mountain Cat (Felis silvestris bieti) and the Arabian Sand Cat (Felis margarita).[19] All the cats in this genus share a common ancestor that probably lived around six million years ago in Asia.[20] Although the exact relationships within the Felidae are still uncertain,[21][22] both the Chinese Mountain Cat and the African Wildcat are close relations of the domestic cat and are both classed as subspecies of the Wildcat Felis silvestris.[4][21] As domestic cats are little altered from wildcats, they can readily interbreed. This hybridization may pose a danger to the genetic distinctiveness of wildcat populations, particularly in Scotland and Hungary.[23]
The domestic cat was first classified as Felis catus by Carolus Linnaeus in the tenth edition of his Systema Naturae of 1758.[2][24] However, due to modern phylogenetics, domestic cats are now usually regarded as another subspecies of the Wildcat Felis silvestris.[4][24][25] This has resulted in mixed usage of the terms, as the domestic cat can be called by its subspecies name, Felis silvestris catus.[1][4] Wildcats have also been referred to as various subspecies of F. catus,[25] but in 2003 the International Commission on Zoological Nomenclature fixed the name for Wildcats as F. silvestris.[26] The most common name in use for the domestic cat remains F. catus, following a convention for domesticated animals of using the earliest (the senior) synonym proposed.[26] Sometimes the domestic cat is called Felis domesticus[27] or Felis domestica,[24] the term coined by German naturalist Johann Christian Polycarp Erxleben in 1777. These are not valid taxonomic names, and Linnaeus' binomial takes precedence.[28]
Cats have either a mutualistic or commensal relationship with humans. However, in comparison to dogs, cats have not undergone major changes during the domestication process, as the form and behavior of the domestic cat are not radically different from those of wildcats, and domestic cats are perfectly capable of surviving in the wild.[29][30] Several natural behaviors and characteristics of Wildcats may have preadapted them for domestication as pets.[30] These traits include their small size, social nature, obvious body language, love of play and relatively high intelligence,[31] they may also have an inborn tendency towards tameness.[30]
There are two main models for how cats were domesticated. In one model, people deliberately tamed cats in a process of artificial selection, as they were useful predators of vermin.[32] However, this model has been criticized as implausible, because there may have been little reward for such an effort: cats do not carry out commands and, although they do eat rodents, other species such as ferrets or terriers may be better at controlling these pests.[4] The alternative idea is that cats were simply tolerated by people and gradually diverged from their 'wild' relatives through natural selection, as they adapted to hunting the vermin found around humans in towns and villages.[4]


Diagram of the general anatomy of a male cat.
Domestic cats are similar in size to the other members of the genus Felis, typically weighing between 4 kilograms (8 lb 13 oz) and 5 kilograms (11 lb 0 oz).[21] However, some breeds, such as the Maine Coon, can exceed 11 kilograms (25 lb). Conversely, very small cats (less than 1.8 kilograms (3 lb 15 oz)) have been reported.[33] The world record for the largest cat is 21.297 kilograms (46 lb 15.2 oz).[34] The smallest adult cat ever officially recorded weighed around 1.36 kilograms (3 lb).[35] Cats average about 23–25 centimeters (9–10 in) in height and 46 centimeters (18.1 in) in head/body length (males being larger than females), with tails averaging 30 centimeters (11.8 in) in length.[36]
Cats have 7 cervical vertebrae like almost all mammals, 13 thoracic vertebrae (humans have 12), 7 lumbar vertebrae (humans have 5), 3 sacral vertebrae like most mammals (humans have 5 because of their bipedal posture), and a variable number of caudal vertebrae in the tail (humans retain 3 to 5 caudal vertebrae, fused into an internal coccyx).[37] The extra lumbar and thoracic vertebrae account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis.[38] Unlike human arms, cat forelimbs are attached to the shoulder by free-floating clavicle bones, which allows them to pass their body through any space into which they can fit their heads.[39]
Skull of a domestic cat.
The cat skull is unusual among mammals in having very large eye sockets and a powerful and specialized jaw.[40] Within the jaw, cats have teeth adapted for killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal neck bite with its two long canine teeth, inserting them between two of the prey's vertebrae and severing its spinal cord, causing irreversible paralysis and death.[41] Compared to other felines, domestic cats have narrowly-spaced canine teeth, which is an adaptation to their preferred prey of small rodents, which have small vertebrae.[41] The premolar and first molar together compose the carnassial pair on each side of the mouth, which efficiently shears meat into small pieces, like a pair of scissors. These are vital in feeding, since cats' small molars cannot chew food effectively.[42]
Cats, like dogs, are digitigrades. They walk directly on their toes, with the bones of their feet making up the lower part of the visible leg.[43] Cats are capable of walking very precisely, because like all felines they directly register; that is, they place each hind paw (almost) directly in the print of the corresponding forepaw, minimizing noise and visible tracks. This also provides sure footing for their hind paws when they navigate rough terrain.
Like almost all members of the Felidae family, cats have protractable claws.[44] In their normal, relaxed position the claws are sheathed with the skin and fur around the toe pads. This keeps the claws sharp by preventing wear from contact with the ground and allows the silent stalking of prey. The claws on the forefeet are typically sharper than those on the hind feet.[45] Cats can voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing, "kneading", or for extra traction on soft surfaces. Most cats have five claws on their front paws, and four on their rear paws.[46] The fifth front claw (the dewclaw) is proximal to the other claws. More proximally, there is a protrusion which appears to be a sixth "finger". This special feature of the front paws, on the inside of the wrists, is the carpal pad, also found on the paws of big cats and dogs. It has no function in normal walking, but is thought to be an anti-skidding device used while jumping. Some breeds of cats are prone to polydactylyism, and may have eight or even ten toes.[46] These are particularly common along the North-East coast of North America.[47]


Cat's eye. Cats have extremely sensitive vision, particularly in dim light.
Cats have excellent night vision and can function at only one-sixth the light level required for human vision.[48] This is partly the result of cat eyes having a tapetum lucidum, which reflects any light that passes through the retina back into the eye, thereby increasing the eye's sensitivity to dim light.[49] Another adaptation to dim light is the large pupils of cats' eyes. Unlike some big cats, such as tigers, domestic cats have slit pupils.[50] These slit pupils can focus bright light without chromatic aberration, and are needed since the domestic cat's pupils are much larger, relative to their eyes, than the pupils of the big cats.[50] Indeed, at low light levels a cat's pupils will expand to cover most of the exposed surface of its eyes.[51] However, domestic cats have rather poor color vision and can only see two colors: blue and green, and are less able to distinguish between red and green,[52] although they can achieve this in some conditions.[53]
Cats' whiskers are highly sensitive to touch.
Cats have excellent hearing and can detect an extremely broad range of frequencies. They can hear higher-pitched sounds than either dogs or humans, detecting frequencies from 55 Hz up to 79 kHz, a range of 10.5 octaves; while humans can only hear from 31 Hz up to 18 kHz, and dogs hear from 67 Hz to 44 kHz, which are both ranges of about 9 octaves.[54][55] Cats do not use this ability to hear ultrasound for communication but it is probably important in hunting,[56] since many species of rodents make ultrasonic calls.[57] Cats' hearing is also extremely sensitive and is among the best of any mammal,[54] being most acute in the range of 500 Hz to 32 kHz.[58] This sensitivity is further enhanced by the cat's large movable outer ears (their pinnae), which both amplify sounds and help a cat sense the direction from which a noise is coming.[56]
Cats have an acute sense of smell, which is due in part to their well-developed olfactory bulb and also to a large surface of olfactory mucosa, in cats this mucosa is about 5.8 cm2 in area, which is about twice that of humans and only 1.7-fold less than the average dog.[59] Cats are very sensitive to pheromones such as 3-Mercapto-3-methylbutan-1-ol,[60] which they use to communicate through urine spraying and marking with scent glands.[61] Cats also respond strongly to plants such as catnip which contains nepetalactone, as they can detect this substance at less than one part per billion.[62] This response is also produced by other plants, such as Silver Vine and Valerian, and may be caused by the smell of these plants mimicking a pheromone and stimulating cats' social or sexual behaviors.[63]
Due to a mutation in an early cat ancestor, one of two genes necessary to taste sweetness may have been lost by the cat family.[64] Their taste buds instead respond to amino acids, bitter tastes and acids.[65] To aid with navigation and sensation, cats have dozens of movable vibrissae (whiskers) over their body, especially their face. These provide information on the width of gaps and on the location of objects in the dark, both by touching objects directly and by sensing air currents; they also trigger protective blink reflexes to protect the eyes from damage.[66]


Blue-eyed cats with white fur have a high incidence of genetic deafness.[67]
The domesticated cat and its closest wild ancestor are both diploid organisms that possess 38 chromosomes[68] and roughly 20,000 genes.[69] About 250 heritable genetic disorders have been identified in cats, many similar to human inborn errors.[70] The high level of similarity among the metabolisms of mammals allows many of these feline diseases to be diagnosed using genetic tests that were originally developed for use in humans, as well as the use of cats in the study of the human diseases.[71][72] An interesting example of a mutation that is shared among all felines, including the big cats, is a mutant chemosensor in their taste buds that prevents them from tasting sweetness, which may explain their indifference to fruits, berries, and other sugary foods.[64] In some breeds of cats congenital deafness is very common, with most white cats (but not albinos) being affected, particularly if they also have blue eyes.[67] The genes responsible for this defect are unknown, but the disease is studied in the hope that it may shed light on the causes of hereditary deafness in humans.[73]
Since a large variety of different coat patterns exist within the various cat breeds, the cat is an excellent animal to study the coat genetics of hair growth and coloration.[74] Several genes interact to produce cats' hair color and coat patterns. Different combinations of these genes give different phenotypes. For example, the enzyme tyrosinase is needed to produce the dark pigment melanin and Burmese cats have a mutant form that is only active at low temperatures, resulting in color appearing only on the cooler ears, tail and paws.[75] A completely inactive gene for tyrosinase is found in albino cats, which therefore lack all pigment.[76] Hair length is determined by the gene for fibroblast growth factor 5, with inactive copies of this gene causing long hair.[77]
The Cat Genome Project, sponsored by the Laboratory of Genomic Diversity at the U.S. National Cancer Institute Frederick Cancer Research and Development Center in Frederick, Maryland, aims to help the development of the cat as an animal model for human hereditary and infectious diseases, as well as contributing to the understanding of the evolution of mammals.[72] This effort led to the publication in 2007 of an initial draft of the genome of a Abyssinian cat called Cinnamon.[69] The existence of a draft genome has led to the discovery of several cat disease genes,[69] and even allowed the development of cat genetic fingerprinting for use in forensics.[78]


Normal physiological values[79]
 Body temperature  38.6°C (101.5°F) 
 Heart rate  120-140 beats per minute 
 Breathing rate  16-40 breaths per minute 
A cat comfortably sleeping with a high skin temperature.
As a familiar and easily-kept animal, the physiology of cats has been particularly well studied and is generally similar to that of other carnivorous mammals.[27] However, several features of cats' physiology are unusual and are probably due to their descent from desert-dwelling species.[27] For instance, cats are able to tolerate quite high temperatures: humans generally start to feel uncomfortable when their skin temperature passes about 44.5 °C (112 °F); in contrast cats show no discomfort until their skin reaches around 52 °C (126 °F),[80] and can tolerate temperatures of up to 56 °C if they have access to water.[81] Cats conserve heat by reducing the flow of blood to their skin and lose heat by evaporation through their mouth. They do not sweat and pant only at very high temperatures.[82] Unusually, a cat's body temperature does not vary throughout the day; this is part of cats' general lack of circadian rhythms and may reflect their tendency to be active at both day and night.[83] As well as being tolerant of high temperatures, cats' feces are usually dry and their urine is also highly concentrated, both of which are adaptations that allow cats to retain as much fluid as possible.[27] Indeed, their kidneys are so efficient that cats can survive on a diet consisting only of meat, with no additional water,[84] and can even rehydrate by drinking seawater.[85][86]
A cat carrying a house mouse.
Cats are obligate carnivores: their physiology has evolved to efficiently process meat, and they have difficulty digesting plant matter.[27] In contrast to omnivores such as rats, which only require about 4% protein in their diet, about 20% of a cat's diet must be protein.[27] Cats are unusually dependent on a constant supply of the amino acid arginine, and a diet lacking arginine causes marked weight loss and can be rapidly fatal.[87] Another unusual feature is that the cat also cannot produce the amino acid taurine, with taurine deficiency causing macular degeneration, where the cat's retina slowly degenerates, causing irreversible blindness.[27] Since cats tend to eat all of their prey, they obtain minerals by digesting animal bones, and a diet composed only of meat may cause calcium deficiency.[27]
A cat's digestive tract is also adapted to meat eating, being much shorter than that of omnivores and having low levels of several of the digestive enzymes that are needed to digest carbohydrates.[88] These traits severely limits the cat's ability to digest and use plant-derived nutrients, as well as certain fatty acids.[88] Despite the cat's meat-oriented physiology, several vegetarian or vegan cat foods have been marketed that are supplemented with chemically synthesized taurine and other nutrients, in attempts to produce a complete diet. However, some of these products still fail to provide all the nutrients that cats require,[89] and diets containing no animal products pose the risk of causing severe nutritional deficiencies.[90]


In captivity, an average life expectancy for male indoor cats at birth is 12 to 14 years,[91] with females usually living a year or two longer. However, there have been records of cats reaching into their 20s and 30s, with the oldest known cat, Creme Puff, dying at a verified age of 38.[92] Having a cat neutered or spayed confers some health benefits, since castrated males cannot develop testicular cancer, spayed females cannot develop uterine or ovarian cancer, and both have a reduced risk of mammary cancer.[93] The lifespan of feral cats is hard to determine accurately, although one study reported a median age of 4.7 years, with a range between 0 to 10 years.[94]


Cats can suffer from a wide range of health problems, including infectious diseases, parasites, injuries and chronic disease. Vaccinations are available for many of these diseases, and domestic cats are regularly given treatments to eliminate parasites such as worms and fleas.


In addition to obvious dangers such as rodenticides, insecticides and weed killers, cats may be poisoned by many chemicals that are usually considered safe.[95] This is because their livers are less effective at some forms of detoxification than those of other animals, including humans and dogs.[27][96] Some of the most common causes of poisoning in cats are antifreeze and rodent baits.[97] It has also been suggested that cats may be particularly sensitive to environmental pollutants.[95][98] When a cat has a sudden or prolonged serious illness without any obvious cause, it is therefore possible that it has been exposed to a toxin.
Human medicines should never be given to cats. .For example, the painkiller paracetamol (also called acetaminophen), sold under brand names such as Tylenol and Panadol is extremely toxic to cats: even very small doses can be fatal and need immediate treatment.^ By on September 19, 2009 at 10:39 am brand for… .
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[99][100] Even aspirin, which is sometimes used to treat arthritis in cats, is much more toxic to them than to humans and must be administered cautiously.[95] Similarly, application of minoxidil (Rogaine) to the skin of cats, either accidentally or by well-meaning owners attempting to counter loss of fur, has sometimes been fatal.[101] Essential oils can be toxic to cats and there have been reported cases of serious illnesses caused by tea tree oil, and tea tree oil-based flea treatments and shampoos.[102]
Other common household substances that should be used with caution around cats include mothballs and other naphthalene products.[95] Phenol-based products are often used for cleaning and disinfecting near cats' feeding areas or litter boxes: such as Pine-Sol, Dettol (Lysol) or hexachlorophene,[95] but these can sometimes be fatal.[103] Ethylene glycol, often used as an automotive antifreeze, is particularly appealing to cats, and as little as a teaspoonful can be fatal.[104] .Some human foods are toxic to cats; for example theobromine in chocolate can cause theobromine poisoning, although few cats will eat chocolate.^ By on November 13, 2009 at 10:52 pm food poisoning symptoms causes… .
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[105] Large amounts of onions or garlic are also poisonous to cats.[95] Many houseplants are also dangerous,[106] such as Philodendron species and the leaves of the Easter Lily, which can cause permanent and life-threatening kidney damage.[107]


Free-ranging cats are active both day and night, although they tend to be slightly more active at night.[108][109] The timing of cats' activity is quite flexible and varied, which means that house cats may be more active in the morning and evening (crepuscular behavior), as a response to greater human activity at these times.[110] House cats have territories that vary quite a lot in size, in one study ranging from seven to 28 hectares.[109] Although they spend the majority of their time in the vicinity of their home, they can range many hundreds of meters from this central point.[109]
An adult Singapura sleeping at the void deck.
Cats conserve energy by sleeping more than most animals, especially as they grow older. The daily duration of sleep varies, usually 12–16 hours, with 13–14 being the average. Some cats can sleep as much as 20 hours in a 24-hour period. The term cat nap refers to the cat's ability to fall asleep (lightly) for a brief period and has entered the English lexicon—someone who nods off for a few minutes is said to be "taking a cat nap". During sleep cats experience short periods of rapid eye movement sleep accompanied by muscle twitches, which suggests that they are dreaming.[111]


Social grooming in a pair of cats.
Although wildcats are solitary, the social behavior of domestic cats is much more variable and ranges from widely-dispersed individuals to feral cat colonies that form around a food source, based on groups of co-operating females.[112][113] Within such groups one cat is usually dominant over the others.[114] Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about ten times larger than those of female cats and may overlap with several females' territories.[61] These territories are marked by urine spraying, by rubbing objects at head height with secretions from facial glands and by defecation.[61] Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling, and if that does not work, by short but noisy and violent attacks. Despite some cats cohabiting in colonies, cats do not have a social survival strategy, or a pack mentality and always hunt alone.[115]
Domestic cats use many vocalizations for communication, including purring, trilling, hissing, growling, snarling and several different forms of meowing.[8] In contrast, feral cats are generally silent.[116] Their types of body language: position of ears and tail, relaxation of whole body, kneading of paws, are all indicators of mood. The tail and ears are particularly important social signal in cats, with a raised tail acting as a friendly greeting.[117][118] Tail raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising their tails less often than subordinate animals.[118] Nose-touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head.[113] However, some pet cats are poorly socialized. In particular older cats may show aggressiveness towards newly-arrived kittens, which may include biting and scratching; this type of behavior is known as Feline Asocial Aggression.[119]
For cats, life in proximity with humans (and other animals kept by humans) amounts to a "symbiotic social adaptation". They may express great affection towards their human companions, especially if they imprint on them at a very young age and are treated with consistent affection. It has been suggested that, ethologically, the human keeper of a cat functions as a sort of surrogate for the cat's mother, and that adult domestic cats live their lives in a kind of extended kittenhood, a form of behavioral neoteny.[120] Conversely, the high-pitched purrs cats make to solicit food may mimic the cries of a hungry human infant, making them particularly hard for humans to ignore.[121]


The hooked papillae on a cat tongue act like a hairbrush to help clean and detangle fur.
Cats are known for their cleanliness, spending many hours licking their coats.[122] The cat's tongue has backwards-facing spines about 500 micrometres long, which are called papillae. These are quite rigid, as they contain keratin.[123] These spines allow cats to groom themselves by licking their fur, with the rows of papillae acting like a hairbrush. Some cats, particularly longhaired cats, occasionally regurgitate hairballs of fur that have collected in their stomachs from grooming. These clumps of fur are usually sausage-shaped and about two to three centimeters long. Hairballs can be prevented with remedies that ease elimination of the hair through the gut, as well as regular grooming of the coat with a comb or stiff brush.[122]


Cats intimidate opponents by arching their backs, raising their fur, turning sideways, and hissing.
With domestic cats, males are more likely to fight than females.[124] With feral cats, the most common reason for cat fighting is when two males are competing to mate with a female: here most fights will be won by the heavier male.[125] Another possible reason for fighting in domestic cats is when the cats have difficulties in establishing a territory within a small home.[124] Female cats will also fight over territory or to defend their kittens. Spaying females and neutering males will decrease or eliminate this behavior in many cases.
When fighting, cats make themselves appear more impressive and threatening by raising their fur and arching their backs, thus increasing their apparent size.[117] Attacks usually comprise powerful slaps to the face and body with the forepaws as well as bites, but serious damage is rare; usually the loser runs away with little more than a few scratches to the face, and sometimes the ears. Cats will also throw themselves to the ground in a defensive posture to rake their opponent's belly with their powerful hind legs.[126]
Normally, serious injuries from fighting will be limited to infections of scratches and bites, though these can occasionally kill cats if untreated. In addition, bites are probably the main route of transmission of feline immunodeficiency virus (FIV).[127] Sexually active males will usually be involved in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to the ears and nose.

Hunting and feeding

Cats feed on small prey, primarily birds and rodents.[128] Feral cats and house cats that are free-fed tend to consume many small meals in a single day, although the frequency and size of meals varies between individuals.[115] Cats use two hunting strategies, either stalking prey actively, or waiting in ambush until an animal comes close enough to be captured. Although it is not certain, the type of strategy used may depend on the prey species in the area, with for example, cats waiting in ambush outside burrows, but tending to actively stalk birds.[129]
Most breeds of cat have a noted fondness for settling in high places, or perching. In the wild, a higher place may serve as a concealed site from which to hunt; domestic cats may strike prey by pouncing from such a perch as a tree branch, as does a leopard.[130] Other possible explanations include that height gives the cat a better observation point, allowing it to survey its territory. During a fall from a high place, a cat can reflexively twist its body and right itself using its acute sense of balance and flexibility.[131] This is known as the cat's "righting reflex". It always rights itself in the same way, provided it has the time to do so, during a fall. The height required for this to occur is around 90 cm (3 feet). Cats without a tail also have this ability, since a cat mostly moves its hind legs and relies on conservation of angular momentum to set up for landing, and the tail is in fact little used for this feat.[132] This leads to the proverb "a cat always lands on its feet."
A domestic cat eating a captured bird.
One poorly understood element of cat hunting behavior is the presentation of prey to human owners. Ethologist Paul Leyhausen proposed that cats adopt humans into their social group, and share excess kill with others in the group according to the local pecking order, in which humans are placed at or near the top.[133] However, anthropologist and animal scientist Desmond Morris, in his 1986 book Catwatching, suggests that when cats bring home mice or birds, they are teaching their human to hunt, or helping their human as if feeding "an elderly cat, or an inept kitten".[134] However, this proposal is inconsistent with the fact that male cats also bring home prey, despite males having no involvement with raising kittens.[129]
Domestic cats select food based on its temperature, smell and texture, strongly disliking chilled foods and responding most strongly to moist foods rich in amino acids, which are similar to meat.[90][115] Cats may reject novel flavors (a response termed neophobia) and learn quickly to avoid foods that have tasted unpleasant in the past.[115] They may also avoid sugary foods and milk; since they are lactose intolerant, these sugars are not easily digested and may cause soft stools or diarrhea.[115][135] They can also develop odd eating habits. Some cats like to eat or chew on other things, most commonly wool, but also plastic, paper, string, or even coal. This condition is called pica and can threaten their health, depending on the amount and toxicity of the items eaten.[136][137]


Domestic cats, especially young kittens, are known for their love of play. This behavior mimics hunting and is important in helping kittens learn to stalk, capture, and kill prey.[138] Cats will also engage in play fighting, with each other and with humans. This behavior may be a way for cats to practice the skills needed for real combat, and might also reduce any fear they associate with launching attacks on other animals.[139]
Due to the close similarity between play and hunting, cats prefer to play with objects that resemble prey, such as small furry toys that move rapidly, but rapidly lose interest (they become habituated) in a toy they have played with before.[140] Cats also tend to play with toys more when they are hungry.[141] String is often used as a toy, but if it is eaten it can become caught at the base of the cat’s tongue and then move into the intestines, a medical emergency which can cause serious illness and death.[142] Due to the risks posed by cats eating string, it is sometimes replaced with a laser pointer's dot, which cats may chase.[143] While concerns have been raised about the safety of these lasers, Professor John Marshall, an ophthalmologist at St Thomas' Hospital, has stated that it would be "virtually impossible" to blind a cat with a laser pointer.[144]


When cats mate, the male tom bites the scruff of the female's neck as she assumes a position conducive to mating.
Female cats are seasonally polyestrous, which means they may have many periods of heat over the course of a year, the season beginning in January or February and ending in late October. Heat periods occur about every two weeks and last about 4 to 7 days.[145] Multiple males will be attracted to a female in heat. The males will fight over her, and the victor wins the right to mate. At first, the female will reject the male, but eventually the female will allow the male to mate. The female will utter a loud yowl as the male pulls out of her. This is because a male cat's penis has a band of about 120-150 backwards-pointing spines, which are about one millimeter long;[146] upon withdrawal of the penis, the spines rake the walls of the female's vagina, which is a trigger for ovulation. After mating, the female will wash her vulva thoroughly. If a male attempts to breed with her at this point, the female will attack him. After about 20 to 30 minutes, once the female is finished grooming, the cycle will repeat.[145]
Because ovulation is not always triggered by a single mating, females may not be impregnated by the first male with which they mate.[147] Furthermore, cats are superfecund; that is, a female may mate with more than one male when she is in heat, with the result that different kittens in a litter may have different fathers.[145]
A kitten with newly-open eyes.
The gestation period for cats is between 64–67 days, with an average length of 66 days.[148] The size of a litter averages three to five kittens, with the first litter usually smaller than subsequent litters. Kittens are weaned at between six and seven weeks, and cats normally reach sexual maturity at 5–10 months (females) and to 5–7 months (males), although this can vary depending on breed.[145] Females can have two to three litters per year, so may produce up to 150 kittens in their breeding span of around ten years.[145]
Cats are ready to go to new homes at about 12 weeks old,[149] or when they are ready to leave their mother. Cats can be surgically sterilized (spayed or castrated) as early as 7 weeks to limit unwanted reproduction.[150] This surgery also prevents undesirable sex-related behavior, such as aggression, territory marking (spraying urine) in males and yowling (calling) in females. Traditionally, this surgery was performed at around six to nine months of age, but it is increasingly being performed prior to puberty, at about three to six months.[151] In the USA approximately 80% of household cats are neutered.[152]



Cats are a cosmopolitan species and are found across much of the world.[29] They are extremely adaptable and are now present on all the continents (including Antarctica) and on 118 of 131 main groups of islands.[153][154] Feral cats can live in forests, grasslands, tundra, coastal areas, agricultural land, scrublands, urban areas and wetlands.[155] Their habitats even include small oceanic islands with no human inhabitants.[156] However, this ability to thrive in almost any terrestrial habitat has led the cat's designation as one of the world's worst invasive species.[157] Despite this general adaptability, the close relatives of domestic cats, the African Wildcat (Felis silvestris lybica) and the Arabian Sand Cat (Felis margarita) both inhabit desert environments,[4] and domestic cats still show similar adaptations and behaviors.[27]

Impact on prey species

Feral cat eating a cottontail rabbit.
To date, there are few scientific data available to assess the impact of cat predation on prey populations. Cat numbers in the UK are growing annually and their abundance is far above the ‘natural’ carrying capacity, due to their population sizes being independent of their prey’s dynamics – i.e. cats are ‘recreational’ hunters.[158] Population densities can be as high as 2000 individuals per km2[159] and the current trend is an increase of 0.5 million cats annually.
Even well-fed domestic cats may hunt and kill, mainly catching small mammals, but also birds, amphibians, reptiles, fish and invertebrates.[128][160] Hunting by domestic cats may be contributing to the decline in the numbers of birds in urban areas, although the importance of this effect remains controversial.[161] In the wild, the introduction of feral cats during human settlement can threaten native species with extinction.[156] In many cases controlling or eliminating the populations of non-native cats can produce a rapid recovery in native animals.[162] However, the ecological role of introduced cats can be more complicated: for example, cats can control the numbers of rats, which also prey on birds' eggs and young, so in some cases eliminating a cat population can actually accelerate the decline of an endangered bird species in the presence of a mesopredator, controlled by cats.[163]
In the Southern Hemisphere, cats are a particular problem in landmasses such as Australasia, where cat species have never been native and there were few equivalent native medium-sized mammalian predators.[164] Native species such as the New Zealand Kakapo and the Australian Bettong, for example, tend to be more ecologically vulnerable and behaviorally "naive" to predation by feral cats.[165] Feral cats have had a major impact on these native species and have played a leading role in the endangerment and extinction of many animals.[166]

Impact on birds

The domestic cat is probably a significant predator of birds. Current UK assessments indicate that they may be accountable for an estimated 64.8 million bird deaths each year.[128] Certain species appear more susceptible than others; for example, 30% of house sparrow mortality is linked to the domestic cat.[167] In the recovery of ringed robins and dunnocks, it was also concluded that 31% of deaths were a result of cat predation.[168]
On islands, birds can contribute as much as 60% of a cat’s diet.[169] In nearly all cases, however, the cat cannot be identified as the sole cause for reducing the numbers of island birds, and in some instances eradication of cats has caused a ‘mesopredator release’ effect;[170] where the suppression of top carnivores creates an abundance of smaller predators that cause a severe decline in their shared prey. Domestic cats are, however, known to be a contributing factor to the decline of many species; a factor that has ultimately led, in some cases, to extinction. The South Island Piopio; Chatham Islands Rail;[168] the Auckland Islands Merganser;[171] and the common diving petrel[172] are a few from a long list, with the most extreme case being the flightless Stephen Island Wren, which was driven to extinction only a few years after its discovery.[173][174]
Some of the same factors that have promoted adaptive radiation of island avifauna over evolutionary time appear to promote vulnerability to non-native species in modern time. The susceptibility inherent of many island birds is undoubtedly due to evolution in the absence of mainland predators, competitors, diseases and parasites. In addition to lower reproductive rates and extended incubation periods.[175] The loss of flight, or reduced flying ability is also characteristic of many island endemics.[176] These biological aspects have increased vulnerability to extinction in the presence of introduced species, such as the domestic cat.[177] Equally, behavioural traits exhibited by island species, such as ‘predatory naivety’[178] and ground-nesting,[175] have also contributed to their susceptibility.

Domesticated cats

Cats are a common companion animal in Europe and North America, and their worldwide population exceeds 500 million.[9] In 1998 there were around 43 million cats in Western Europe, 33 million in Eastern Europe, seven million in Japan and three million in Australia.[179] A 2007 report stated that about 37 million US households owned cats, with an average of 2.2 cats per household giving a total population of around 82 million.[180] This is slightly more than the 72 million pet dogs in that country.[180] Although cat ownership has commonly been associated with women,[181] a 2007 Gallup poll reported that men and women were equally likely to own a cat.[182] The ratio of pedigree/purebred cats to random-bred cats varies from country to country. However, generally speaking, purebreds are less than ten percent of the total population.[183]
A cat pelt.
According to the Humane Society of the United States, as well as being kept as pets, cats are also used in the international fur trade.[184] Cat fur is used in coats, gloves, hats, shoes, blankets and stuffed toys. About 24 cats are needed to make a cat fur coat.[185] This use has now been outlawed in several countries, including the United States, Australia and the European Union.[186] However, some cat furs are still made into blankets in Switzerland as folk remedies that are believed to help rheumatism.[187]
It has long been common for cats to be eaten in some parts of Italy, China and in some other Asian countries. According to the Chengdu Business Daily, people in southern China's Guangdong province ate 10,000 cats a day.[188] Animal People estimates that 4 million cats are killed and consumed in Asia every year.[189] Cat stew is also a Tuscan (Italian) specialty.[190]

Domesticated varieties

The concept of a cat breed appeared in Britain during the late 19th century.[191] The current list of cat breeds is quite large: with the Cat Fanciers' Association recognizing 41 breeds, of which 16 are "natural breeds" that probably emerged before humans began breeding pedigree cats, while the others were developed over the latter half of the 20th century.[29] The owners and breeders of show cats compete to see whose animal bears the closest resemblance to the "ideal" definition and standard of the breed (see selective breeding). Because of common crossbreeding in populated areas, many cats are simply identified as belonging to the homogeneous breeds of domestic longhair and domestic shorthair, depending on their type of fur. In the United Kingdom and Australasia, non-purebred cats are referred in slang as moggies (derived from "Maggie", short for Margaret, reputed to have been a common name for cows and calves in 18th century England and latter applied to housecats during the Victorian era).[192] In the United States, a non-purebred cat is sometimes referred to in slang as a barn or alley cat, even if it is not a stray. Cats come in a variety of colors and patterns. These are physical properties and should not be confused with a breed of cat. Some original cat breeds that have a distinct phenotype that is the main type occurring naturally as the dominant domesticated cat type in their region of origin are sometimes considered as subspecies and also have received names as such in nomenclature, although this is not supported by feline biologists. Some of these cat breeds are:
  • F. catus anura - the Manx
  • F. catus siamensis - the Siamese
  • F. catus cartusenensis - the Chartreux
  • F. catus angorensis - the Turkish Angora

Coat patterns

Cat coat genetics can produce a variety of coat patterns. Some of the most common are:
Bicolor, Tuxedo and Van 
This pattern varies between the tuxedo cat which is mostly black with a white chest, and possibly markings on the face and paws/legs, all the way to the Van pattern (so named after the Lake Van area in Turkey, which gave rise to the Turkish Van breed), where the only colored parts of the cat are the tail (usually including the base of the tail proper), and the top of the head (often including the ears). There are several other terms for amounts of white between these two extremes, such as Harlequin or jellicle cat. Bicolor cats can have as their primary (non-white) color black, red, any dilution thereof, and tortoiseshell (see below for definition).
Mackerel tabby cat, showing the characteristic "M" on its forehead.
A Chilean torbie cat.
Tabby cat 
Striped, with a variety of patterns. The classic blotched tabby (or marbled) pattern is the most common and consists of butterflies and bullseyes. The mackerel or striped tabby is a series of vertical stripes down the cat's side (resembling the fish). This pattern broken into spots is referred to as a spotted tabby. Finally, the tabby markings may look like a series of ticks on the fur, thus the ticked tabby, which is almost exclusively associated with the Abyssinian breed of cats. The worldwide evolution of the cat means that certain types of tabby are associated with certain countries; for instance, blotched tabbies are quite rare outside NW Europe, where they are the most common type.
Female tortoiseshell-and-white cat.
Tortoiseshell and Calico.
This cat is also known as a Calimanco cat or Clouded Tiger cat, and by the nickname "tortie." In the cat fancy, a tortoiseshell cat is randomly patched over with red (or its dilute form, cream) and black (or its dilute blue) mottled throughout the coat. Additionally, the cat may have white spots in its fur, which make it a "tortoiseshell and white" cat or, if there is a significant amount of white in the fur and the red and black colors form a patchwork rather than a mottled aspect, the cat will be called a calico. All calicos are tortoiseshell (as they carry both black and red), but not all tortoiseshells are calicos (which requires a significant amount of white in the fur and patching rather than mottling of the colors). The calico is also sometimes called a tricolor cat. The Japanese refer to this pattern as mi-ke (meaning "triple fur"), while the Dutch call these cats lapjeskat (meaning "patches cat"). A true tricolor must consist of three colors: a reddish color, dark or light; white; and one other color, typically a brown, black, or blue.[193] Both tortoiseshell and calico cats are typically female because the coat pattern is the result of differential X chromosome inactivation in females (which, as with all normal female mammals, have two X chromosomes). Conversely, cats where the overall color is ginger (orange) are commonly male (roughly in a 3:1 ratio). In a litter sired by a ginger tom, the females will be tortoiseshell or ginger. Male tortoiseshells can occur as a result of chromosomal abnormalities (often linked to sterility) or by a phenomenon known as mosaicism, where two early stage embryos are merged into a single kitten.
Siamese cat, classical colorpoint pattern.
The colorpoint pattern is most commonly associated with Siamese cats, but may also appear in any domesticated cat. A colorpointed cat has dark colors on the face, ears, feet, and tail, with a lighter version of the same color on the rest of the body, and possibly some white. .The exact name of the colorpoint pattern depends on the actual color, so there are seal points (dark brown), chocolate points (warm lighter brown), blue points (dark gray), lilac or frost points (silvery gray-pink), red or flame points (orange), and tortie (tortoiseshell mottling) points, among others.^ By on October 9, 2009 at 2:47 pm red orange blue green yellow… .
  • Stay Dougie » Mos Def In Your Neighborhood Boneyard With Stussy 11 January 2010 15:14 UTC [Source type: FILTERED WITH BAYES]

This pattern is the result of a temperature sensitive mutation in one of the enzymes in the metabolic pathway from tyrosine to pigment, such as melanin; thus, little or no pigment is produced except in the extremities or points where the skin is slightly cooler. For this reason, colorpointed cats tend to darken with age as bodily temperature drops; also, the fur over a significant injury may sometimes darken or lighten as a result of temperature change.
The tyrosine pathway also produces neurotransmitters, thus mutations in the early parts of that pathway may affect not only pigment, but also neurological development. This results in a higher frequency of cross-eyes among colorpointed cats, as well as the high frequency of cross-eyes in white tigers.
White cats
True albinism (a mutation of the tyrosinase gene) is quite rare in cats. Much more common is the appearance of white coat color due to a lack of melanocytes in the skin. A higher frequency of deafness in white cats is due to a reduction in the population and survival of melanoblast stem cells, which in addition to creating pigment producing cells, develop into a variety of neurological cell types. White cats with one or two blue eyes have a particularly high likelihood of being deaf.
Smoke cats
The bottom eighth of each hair is white or creamy-white, with the rest of the hair being a solid color. Genetically this color is a non-agouti cat with the dominant inhibitor gene; a non-agouti version of the silver tabby. Smoke cats will look solid colored until they move, when the white undercoat becomes apparent. It is mostly found in pedigreed cats (especially longhair breeds) but also present in some domestic longhaired cats.

Body types

Cats can also come in several body types, ranging between two extremes:
Not a specific breed, but any cat with an elongated slender build, almond-shaped eyes, long nose, large ears (the Siamese and Oriental Shorthair breeds are examples of this).
less slender than the oriental type, but nevertheless a cat with a slight build and generally athletic look. Typical example breeds would be the Abyssinian cat and the Turkish Angora. Some people consider the foreign and oriental body types as being the same, however.
More or less the middle range of body conformation types, this type of cat is less slender without being stocky. Example breeds would be the Devon Rex and the Egyptian Mau.
these cats look more rounded without looking too stocky. Example breeds would be the American Shorthair and British Shorthair.
Any cat with a short, muscular, compact build, roundish eyes, short nose, and small ears. Persian cats and Exotic cats are two prime examples of such a body type.

Effects on human health

Because of their small size, domesticated house cats pose little physical danger to adult humans. However, in the USA cats inflict about 400,000 bites per year, with 90% of these bites coming from provoked animals; this number represents about one in ten of all animal bites.[194] Many cat bites will become infected,[195] sometimes with serious consequences such as cat-scratch disease, or, more rarely, rabies.[194] Cats may also pose a danger to pregnant women and immunosuppressed individuals, since their feces can transmit toxoplasmosis.[196] A large percentage of cats are infected with this parasite, with infection rates ranging from around 40 to 60% in both domestic and stray cats worldwide.[197][198][199]
Allergic reactions to cat dander and/or cat saliva are common.[200] Some humans who are allergic to cats—typically manifested by hay fever, asthma, or a skin rash—quickly acclimate themselves to a particular animal and live comfortably in the same house with it, while retaining an allergy to cats in general.[201] Whether the risk of developing allergic diseases such as asthma is increased or decreased by cat ownership is uncertain.[202][203] Some owners cope with this problem by taking allergy medicine, along with bathing their cats frequently, since weekly bathing will reduce the amount of dander shed by a cat.[204] There have also been attempts to breed hypoallergenic cats, which would be less likely to provoke an allergic reaction.[205]
As well as posing health risks, interactions with cats may improve health and reduce physical responses to stress: for example the presence of cats may moderate increased blood pressure.[206] Cat ownership may also improve psychological health by providing emotional support and dispelling feelings of depression, anxiety and loneliness.[207] Indeed, their ability to provide companionship and friendship are common reasons given for owning a cat.[182]

Indoor scratching

A natural behavior in cats is to periodically hook their front claws into suitable surfaces and pull backwards. This marks territory, exercises their legs, as well as cleaning and sharpening their claws.[208] Indoor cats benefit from being provided with a scratching post so that they are less likely to use carpet or furniture which they can easily ruin.[209] Commercial scratching posts typically are covered in carpeting or upholstery, but some authorities advise against this practice, as not making it clear to the cat which surfaces are permissible and which are not; they suggest using a plain wooden surface, or reversing the carpeting on the posts so that the rougher texture of the carpet backing is a more attractive alternative to the cat than the floor covering. Scratching posts made of sisal rope or corrugated cardboard are also common.
Close-up of a cat's claw, with the quick clearly visible.
Although scratching can serve cats to keep their claws from growing excessively long, their nails can be trimmed if necessary. Another response to indoor scratching is onychectomy, commonly known as declawing. This is a surgical procedure to remove the claw and first bone of each digit of a cat's paws. Declawing is most commonly only performed on the front feet. A related procedure is tendonectomy, which involves cutting a tendon needed for cats to extend their claws.[210] Declawing is a major surgical procedure and can produce pain, infections and permanent lameness.[210] Since this surgery is not performed for the benefit of the animal, it is controversial and remains uncommon outside of North America.[211] In many countries, it is prohibited by animal welfare laws. Although widely practiced in the US, declawing is ethically controversial within the American veterinary community.[212] Both the Humane Society of the United States and the American Society for the Prevention of Cruelty to Animals strongly discourage or condemn the procedure.[213] A declawed cat that is lost or abandoned lacks any chance of successful hunting and is rendered unable to climb trees to safety and escape predators like stray dogs, a combination that essentially gives it no chances of survival. Since cats use claws for traction when running or to prevent skidding, loss of claws or use thereof decreases their mobility, too. Also, a declawed is unable to defend itself in a fight, whether against other pets at home or predators and other ferals if it ends up a stray.


Toilet-trained house cat.
Cats bury their urine and feces. Indoor cats are usually provided with a litter box containing litter, typically bentonite, but sometimes other absorbent material such as shredded paper or wood chips, or sometimes sand or similar material. It should be cleaned daily and changed often, depending on the number of cats in a household and the type of litter; if it is not kept clean, a cat may be fastidious enough to find other locations in the house for urination or defecation. This may also happen for other reasons; for instance, if a cat becomes constipated and defecation is uncomfortable, it may associate the discomfort with the litter box and avoid it in favor of another location.
Daily attention to the litter box also serves as a monitor of the cat's health. Bentonite or clumping litter is a variation which absorbs urine into clumps which can be sifted out along with feces, and thus stays cleaner longer with regular sifting, but has sometimes been reported to cause health problems in some cats.[214] Those with toxoplasmosis-infected cats living in habitat areas of sea otters[215] may wish to dispose of droppings in the trash, rather than flushing them down the toilet.[216] Some cats can be trained to use the human toilet, eliminating the litter box and its attendant expense, unpleasant odor, and the need to use landfill space for disposal. Training may involve four to six weeks of incremental moves, such as moving and elevating the litter box until it is near the toilet, as well as employing an adapter such as a bowl or small box to suspend the litter above the toilet bowl.[217] When training is complete, the cat uses the toilet by squatting on the toilet seat over the bowl.

Feral cats

American feral farm cat.
Feral cats are wild cats that are unfamiliar with humans and roam freely in urban or rural areas.[218] The numbers of feral cats are not known, but estimates of the US feral population range from 25 to 60 million.[218] Feral cats may live alone, but most are found in large groups called feral colonies, which occupy a specific territory and are usually associated with a source of food.[219] Famous feral cat colonies are found in Rome around the Colosseum and Forum Romanum, with cats at some of these sites being fed and vetted by volunteers.[220]
Public attitudes towards feral cats vary widely: ranging from seeing them as free-ranging pets, to regarding them as vermin.[221] One common approach to reducing the feral cat population is termed trap-neuter-return, where the cats are trapped, neutered, immunized against rabies and the feline leukemia virus, and then released. Before releasing them back into their feral colonies, the attending veterinarian often nips the tip off one ear to mark the feral as neutered and inoculated, since these cats may be trapped again.[219] Volunteers continue to feed and give care to these cats throughout their lives, and not only is their lifespan greatly increased, but behavior and nuisance problems, due to competition for food, are also greatly reduced.[219]

History and mythology

Egyptian sculpture at the Louvre.
Traditionally, historians tended to think that ancient Egypt was the site of cat domestication, due to the clear depictions of house cats in Egyptian paintings about 3,600 years old.[4] However, in 2004, a Neolithic grave was excavated in Shillourokambos, Cyprus that contained the skeletons, laid close to one another, of both a human and a cat. The grave is estimated to be 9,500 years old, pushing back the earliest known feline-human association significantly.[6][222][223] The cat specimen is large and closely resembles the African wildcat (Felis silvestris lybica), rather than present-day domestic cats. This discovery, combined with genetic studies, suggest that cats were probably domesticated in Cyprus and the Near East, in the Fertile Crescent around the time of the development of agriculture.[4]
In ancient Egypt cats were sacred animals, with the Bast often depicted in cat form, sometimes taking on the warlike aspect of a lioness.[224] The Romans are often credited with introducing the domestic cat from Egypt to Europe.[225] However, it is possible that cats were already kept in Europe prior to the Roman Empire, as they may have already been present in Britain in the late Iron Age.[32] Domestic cats were spread throughout much of the rest of the world during the Age of Discovery, as they were carried on sailing ships to control shipboard rodents and as good-luck charms.[225]
Freya and her cats.
Several ancient religions believed that cats are exalted souls, companions or guides for humans, that they are all-knowing but are mute so they cannot influence decisions made by humans. In Japan, the Maneki Neko is a cat that is a symbol of good fortune. Although there are no sacred species in Islam, some writers have stated that Muhammad had a favorite cat, Muezza.[226] He is reported to have loved cats so much that "he would do without his cloak rather than disturb one that was sleeping on it".[227]
Freyja—the goddess of love, beauty, and fertility in Norse mythology—is depicted as riding a chariot drawn by cats.
Many cultures have negative superstitions about cats. An example would be the belief that a black cat "crossing your path" leads to bad luck, or that cats are witches' familiars used to augment a witch's powers and skills. This led to the widespread extermination of cats in Europe in medieval times. The Black Plague was spread by fleas carried by infected rats, and the killing of cats ostensibly caused an increase in the rat population. The killing of cats in Medieval Ypres is commemorated in the innocuous present-day Kattenstoet (cat parade).
According to a myth in many cultures, cats have nine (or sometimes seven) lives. The myth is attributed to the natural suppleness and swiftness cats exhibit to escape life-threatening situations.[228] Also lending credence to this myth is that falling cats often land on their feet because of an inbuilt automatic twisting reaction and are able to twist their bodies around to land feet first, though they can still be injured or killed by a high fall.[229]

See also


  1. ^ a b c Wozencraft, W. C. (16 November 2005). Wilson, D. E., and Reeder, D. M. (eds). ed. Mammal Species of the World (3rd edition ed.). Johns Hopkins University Press. ISBN 0-801-88221-4. 
  2. ^ a b Linnaeus, Carolus (1766) [1758] (in (Latin)). Systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. 1 (12th ed.). Holmiae (Laurentii Salvii). p. 62. Retrieved 2008-04-02. 
  3. ^ ITIS. "ITIS Standard Report Page: Felis catus domestica". 
  4. ^ a b c d e f g h i j Driscoll CA, Macdonald DW, O'Brien SJ, CA (June 2009). "In the Light of Evolution III: Two Centuries of Darwin Sackler Colloquium: From wild animals to domestic pets, an evolutionary view of domestication". Proc. Natl. Acad. Sci. U.S.A. 106 (S1): 9971–9978. doi:10.1073/pnas.0901586106. ISSN 0027-8424. PMID 19528637. 
  5. ^ "Housecat" in the American Heritage Dictionary
  6. ^ a b "Oldest Known Pet Cat? 9500-Year-Old Burial Found on Cyprus". National Geographic News. 2004-04-08. Retrieved 2007-03-06. 
  7. ^ Carlos A. Driscoll, Juliet Clutton-Brock, Andrew C. Kitchener and Stephen J. O'Brien. "The Evolution of House Cats". Scientific American. Retrieved 26 August 2009. 
  8. ^ a b Moelk, Mildred (1944-04). "Vocalizing in the House-Cat; A Phonetic and Functional Study". The American Journal of Psychology 57 (2): 184-205. ISSN 00029556. Retrieved 2010-03-04. 
  9. ^ a b Wade, Nicholas (2007-06-29). "Study Traces Cat's Ancestry to Middle East". The New York Times. Retrieved 2008-04-02. 
  10. ^ "Meet Helen and Aphrodite, Cyprus's indigenous cats". China Daily. Retrieved 2009-11-03. 
  11. ^ "Cat". The Online etymology dictionary. Retrieved 2007-05-15. 
  12. ^ Crum, Walter Ewing. A Coptic Dictionary]. Oxford: Clarendon Press, 1939: 601
  13. ^ "Le chat: origines et étymologie." Chat et compagnie. 2006
  14. ^ SenenAnep Meritamen. "English to Egyptian Dictionary." posted August 29, 2004. Ancient Worlds. AncientWorlds LLC, 2002.
  15. ^ Webster's Encyclopedic Unabridged Dictionary of the English Language. New York: Gramercy Books, 1996: 1571.
  16. ^
  17. ^ Johnson, Warren; O’brien, Stephen J. (1997-01-01). "Phylogenetic reconstruction of the felidae using 16S rRNA and NADH-5 mitochondrial genes". Journal of Molecular Evolution 44 (0): S98–S116. doi:10.1007/PL00000060. 
  18. ^ "ITIS Standard Report Page: Felis". Integrated Taxonomic Information System. Retrieved 26 August 2009. 
  19. ^ Stefoff, Rebecca (2003-11). Cats. Benchmark Books (NY). p. 34. ISBN 0761415777. 
  20. ^ Johnson, WE; Eizirik, E; Pecon-Slattery, J; Murphy, WJ; Antunes, A; Teeling, E; O'brien, SJ (2006). "The Late Miocene Radiation of Modern Felidae: A Genetic Assessment". Science 311 (5757): 73–77. doi:10.1126/science.1122277. PMID 16400146. Retrieved 2009-09-30. 
  21. ^ a b c Mattern, Michelle Y.; McLennan, Deborah A. (2000). "Phylogeny and Speciation of Felids". Cladistics 16 (2): 232–253. doi:10.1111/j.1096-0031.2000.tb00354.x. 
  22. ^ Masuda, R; Lopez, JV; Slattery, JP; Yuhki, N; O'brien, SJ (1996-12). "Molecular Phylogeny of Mitochondrial Cytochrome b and 12S rRNA Sequences in the Felidae: Ocelot and Domestic Cat Lineages". Molecular Phylogenetics and Evolution 6 (3): 351–365. doi:10.1006/mpev.1996.0085. ISSN 1055-7903. PMID 8975691. Retrieved 2009-09-30. 
  23. ^ Oliveira R, Godinho R, Randi E, Alves PC, R (September 2008). "Hybridization versus conservation: are domestic cats threatening the genetic integrity of wildcats (Felis silvestris silvestris) in Iberian Peninsula?". Philos. Trans. R. Soc. Lond., B, Biol. Sci. 363 (1505): 2953–61. doi:10.1098/rstb.2008.0052. ISSN 0962-8436. PMID 18522917. 
  24. ^ a b c Wozencraft, W. C. (16 November 2005). Wilson, D. E., and Reeder, D. M. (eds). ed. Mammal Species of the World (3rd edition ed.). Johns Hopkins University Press. pp. 534–535. ISBN 0-801-88221-4. 
  25. ^ a b Wozencraft, W. C. (16 November 2005). Wilson, D. E., and Reeder, D. M. (eds). ed. Mammal Species of the World (3rd edition ed.). Johns Hopkins University Press. pp. 536–537. ISBN 0-801-88221-4. 
  26. ^ a b ICZN (2003-03-31). "Opinion 2027". Bulletin of Zoological Nomenclature (International Commission on Zoological Nomenclature) 60. Retrieved 2008-04-02. 
  27. ^ a b c d e f g h i j MacDonald ML, Rogers QR, Morris JG, ML (1984). "Nutrition of the domestic cat, a mammalian carnivore". Annu. Rev. Nutr. 4: 521–62. doi:10.1146/ ISSN 0199-9885. PMID 6380542. 
  28. ^ Vella, Carolyn M.; Lorraine M. Shelton, John J. McGonagle, Terry W. Stanglein (1999). Robinson's Genetics for Cat Breeders and Veterinarians (4 ed.). Butterworth-Heinemann. p. 3. ISBN 0750640693. 
  29. ^ a b c Lipinski, MJ; Froenicke, L; Baysac, KC; Billings, NC; Leutenegger, CM; Levy, AM; Longeri, M; Niini, T et al. (2008-01). "The ascent of cat breeds: Genetic evaluations of breeds and worldwide random-bred populations". Genomics 91 (1): 12–21. doi:10.1016/j.ygeno.2007.10.009. ISSN 0888-7543. PMID 18060738. PMC 2267438. Retrieved 2009-08-28. 
  30. ^ a b c Cameron-Beaumont, Charlotte; Lowe, Sarah E.; Bradshaw, John W. S. (2002). "Evidence suggesting preadaptation to domestication throughout the small Felidae" (PDF). Biological Journal of the Linnean Society 75 (3): 361–366. doi:10.1046/j.1095-8312.2002.00028.x. Retrieved 2009-09-29. 
  31. ^ (Fogle 1981, pp. 12–17)
  32. ^ a b O'Connor, T. P. (2007). "Wild or domestic? Biometric variation in the cat Felis silvestris Schreber". International Journal of Osteoarchaeology 17 (6): 581–595. doi:10.1002/oa.913. 
  33. ^ "Dwarf, Midget and Miniature cats". Retrieved 2007-03-06. 
  34. ^ Cat World (2008). Cat World Records: Heaviest Cat. Retrieved on 2008-07-30 from
  35. ^ retrieved on 2008/29/11
  36. ^ "Domestic Cat". SeaWorld. Retrieved 2009-03-04. 
  37. ^ (Walker 1982, p. 11)
  38. ^ (Walker 1982, p. 16)
  39. ^ "Cat Skeleton". Archived from the original on 2006-12-06. Retrieved 2006-12-12. 
  40. ^ (Case 2003, p. 35)
  41. ^ a b Smith, Patricia; Eitan Tchernov (1992). Structure, function and evolution of teeth. Freund Publishing House Ltd.. p. 217. ISBN 9652222704, 9789652222701. 
  42. ^ (Case 2003, p. 37)
  43. ^ Lacquaniti F, Grasso R, Zago M, F (1 August 1999). "Motor Patterns in Walking". News Physiol. Sci. 14 (4): 168–174. ISSN 0886-1714. PMID 11390844. 
  44. ^ Russell, Anthony P.; Bryant, Harold N. (2001). "Claw Retraction and Protraction in the Carnivora: The Cheetah (Acinonyx Jubatus) as an Atypical Felid". Journal of Zoology 254 (01): 67–76. doi:10.1017/S0952836901000565. Retrieved 2009-08-28. 
  45. ^ Armes, Annetta F. (1900-12-22). "Outline of Cat Lessons". The School Journal LXI: 659. Retrieved 2007-11-12. 
  46. ^ a b Danforth CH. (1947). "Heredity of Polydactyly in the Cat" (PDF). J. Hered. 38: 107–112. 
  47. ^ Lettice, LA; Hill, AE; Devenney, PS; Hill, RE (2008-04-01). "Point mutations in a distant sonic hedgehog cis-regulator generate a variable regulatory output responsible for preaxial polydactyly". Human Molecular Genetics 17 (7): 978–985. doi:10.1093/hmg/ddm370. ISSN 1460-2083. PMID 18156157. 
  48. ^ (Case 2003, p. 43)
  49. ^ Ollivier, F. J.; Samuelson, DA; Brooks, DE; Lewis, PA; Kallberg, ME; Komáromy, AM (2004). "Comparative morphology of the tapetum lucidum (among selected species)". Veterinary Ophthalmology 7 (1): 11–22. doi:10.1111/j.1463-5224.2004.00318.x. PMID 14738502. 
  50. ^ a b Malmström T, Kröger RH, T (January 2006). "Pupil shapes and lens optics in the eyes of terrestrial vertebrates" (Free full text). J. Exp. Biol. 209 (Pt 1): 18–25. doi:10.1242/jeb.01959. ISSN 0022-0949. PMID 16354774. 
  51. ^ Hammond, P.; Mouat, G.S.V. (1985). "The relationship between feline pupil size and luminance". Experimental Brain Research 59 (3): 485–490. doi:10.1007/BF00261338. 
  52. ^ Loop MS, Bruce LL, MS (March 1978). "Cat color vision: the effect of stimulus size". Science 199 (4334): 1221–2. doi:10.1126/science.628838. ISSN 0036-8075. PMID 628838. 
  53. ^ Schneider H, Beller FK (1 April 1993). "The Spectral Sensitivity of Dark- and Light-adapted Cat Retinal Ganglion Cells". Journal of Neuroscience 13 (4): 1543–1550. PMID 8463834. 
  54. ^ a b Heffner, RS (2004-11). "Primate hearing from a mammalian perspective" (PDF). The Anatomical Record. Part A, Discoveries in Molecular, Cellular, and Evolutionary Biology 281 (1): 1111–1122. doi:10.1002/ar.a.20117. ISSN 1552-4884. PMID 15472899. Retrieved 2009-08-20. 
  55. ^ Heffner, Henry E. (1998-05). "Auditory awareness". Applied Animal Behaviour Science 57 (3-4): 259–268. doi:10.1016/S0168-1591(98)00101-4. ISSN 0168-1591. Retrieved 2009-08-20. 
  56. ^ a b (Sunquist & Sunquist 2002, p. 10)
  57. ^ Blumberg, M S (December 1992). "Rodent ultrasonic short calls: locomotion, biomechanics, and communication". Journal of Comparative Psychology 106 (4): 360–365. doi:10.1037/0735-7036.106.4.360. ISSN 0735-7036. PMID 1451418. 
  58. ^ Heffner, R S; Heffner, HE (1985). "Hearing range of the domestic cat" (PDF). Hearing Research 19 (1): 85–88. doi:10.1016/0378-5955(85)90100-5. ISSN 0378-5955. PMID 4066516. Retrieved 2009-08-20. 
  59. ^ Moulton, David G. (1967-08-01). "Olfaction in Mammals". Amer. Zool. 7 (3): 421–429. doi:10.1093/icb/7.3.421. Retrieved 2009-08-20. 
  60. ^ M. Miyazaki, T. Yamashita, Y. Suzuki, Y. Saito, S. Soeta, H. Taira, and A. Suzuki, M (October 2006). "A major urinary protein of the domestic cat regulates the production of felinine, a putative pheromone precursor" (PDF). Chem. Biol. 13 (10): 1071–1079. doi:10.1016/j.chembiol.2006.08.013. ISSN 1074-5521. PMID 17052611. 
  61. ^ a b c Sommerville, B. A. (1998). "Olfactory awareness". Applied Animal Behaviour Science 57 (3-4): 269–286. doi:10.1016/S0168-1591(98)00102-6. ISSN 0168-1591. 
  62. ^ Grognet, Jeff (1990-06). "Catnip: Its uses and effects, past and present". The Canadian Veterinary Journal 31 (6): 455–456. 
  63. ^ Tucker, Arthur; Sharon Tucker (1988). "Catnip and the catnip response". Economic Botany 42 (2): 214-231. doi:10.1007/BF02858923. 
  64. ^ a b Li, X; Li, W; Wang, H; Cao, J; Maehashi, K; Huang, L; Bachmanov, AA; Reed, DR et al. (July 2005). "Pseudogenization of a Sweet-Receptor Gene Accounts for Cats' Indifference toward Sugar" (Free full text). PLOS Genetics (Public Library of Science) 1 (1): e3. doi:10.1371/journal.pgen.0010003. ISSN 1553-7390. PMID 16103917. PMC 1183522. Retrieved 2006-11-08. 
  65. ^ Bradshaw, John W. S. (2006-07-01). "The Evolutionary Basis for the Feeding Behavior of Domestic Dogs (Canis familiaris) and Cats (Felis catus)". J. Nutr. 136 (7): 1927S–1931. ISSN 0022-3166. PMID 16772461. Retrieved 2009-08-25. 
  66. ^ (Case 2003, p. 47)
  67. ^ a b Strain GM, GM (January 1996). "Aetiology, prevalence and diagnosis of deafness in dogs and cats". Br. Vet. J. 152 (1): 17–36. doi:10.1016/S0007-1935(96)80083-2. ISSN 0007-1935. PMID 8634862. 
  68. ^ Nie W, Wang J, O'Brien PC, W (2002). "The genome phylogeny of domestic cat, red panda and five mustelid species revealed by comparative chromosome painting and G-banding". Chromosome Res. 10 (3): 209–22. doi:10.1023/A:1015292005631. ISSN 0967-3849. PMID 12067210. 
  69. ^ a b c Pontius JU, Mullikin JC, Smith DR, JU (November 2007). "Initial sequence and comparative analysis of the cat genome" (Free full text). Genome Res. 17 (11): 1675–89. doi:10.1101/gr.6380007. ISSN 1088-9051. PMID 17975172. PMC 2045150. 
  70. ^ O'Brien SJ, Johnson W, Driscoll C, Pontius J, Pecon-Slattery J, Menotti-Raymond M, SJ (June 2008). "State of cat genomics". Trends Genet. 24 (6): 268–79. doi:10.1016/j.tig.2008.03.004. ISSN 0168-9525. PMID 18471926. 
  71. ^ Sewell AC, Haskins ME, Giger U, AC (September 2007). "Inherited metabolic disease in companion animals: searching for nature's mistakes". Vet. J. 174 (2): 252–9. doi:10.1016/j.tvjl.2006.08.017. ISSN 1090-0233. PMID 17085062. 
  72. ^ a b O'Brien SJ, Menotti-Raymond M, Murphy WJ, Yuhki N, SJ (2002). "The Feline Genome Project". Annu. Rev. Genet. 36: 657–86. doi:10.1146/annurev.genet.36.060602.145553. ISSN 0066-4197. PMID 12359739. 
  73. ^ Saada AA, Niparko JK, Ryugo DK, AA (October 1996). "Morphological changes in the cochlear nucleus of congenitally deaf white cats". Brain Res. 736 (1-2): 315–28. doi:10.1016/0006-8993(96)00719-6. ISSN 0006-8993. PMID 8930338. 
  74. ^ Robinson, Roy; Vella, Carolyn M.; Lorraine Shelton; McGonagle, John J.; Carolyne Vella (1999). Robinson's genetics for cat breeders and veterinarians. Oxford: Butterworth-Heinemann. ISBN 0-7506-4069-3. 
  75. ^ Lyons LA, Imes DL, Rah HC, Grahn RA, LA (April 2005). "Tyrosinase mutations associated with Siamese and Burmese patterns in the domestic cat (Felis catus)". Anim. Genet. 36 (2): 119–26. doi:10.1111/j.1365-2052.2005.01253.x. ISSN 0268-9146. PMID 15771720. 
  76. ^ Imes DL, Geary LA, Grahn RA, Lyons LA, DL (April 2006). "Albinism in the domestic cat (Felis catus) is associated with a tyrosinase (TYR) mutation" (Free full text). Anim. Genet. 37 (2): 175–8. doi:10.1111/j.1365-2052.2005.01409.x. ISSN 0268-9146. PMID 16573534. 
  77. ^ Kehler JS, David VA, Schäffer AA, JS (September 2007). "Four independent mutations in the feline fibroblast growth factor 5 gene determine the long-haired phenotype in domestic cats" (Free full text). J. Hered. 98 (6): 555–66. doi:10.1093/jhered/esm072. ISSN 0022-1503. PMID 17767004. 
  78. ^ Menotti-Raymond M, David VA, Stephens JC, Lyons LA, O'Brien SJ, M (November 1997). "Genetic individualization of domestic cats using feline STR loci for forensic applications" (Free full text). J. Forensic Sci. 42 (6): 1039–51. ISSN 0022-1198. PMID 9397545. 
  79. ^ (Kahn & Line 2007, p. 330)
  80. ^ (Case 2003, p. 46)
  81. ^ Subcommittee on Dog and Cat Nutrition Co (2006). Nutrient requirements of dogs and cats. Washington, D.C: National Academies Press. pp. 292. ISBN 0-309-08628-0. 
  82. ^ Adams T, Morgan ML, Hunter WS, Holmes KR (December 1970). "Temperature regulation of the unanesthetized cat during mild cold and severe heat stress". J Appl Physiol 29 (6): 852–8. PMID 5485356. 
  83. ^ (Committee on Animal Nutrition 1986, p. 1)
  84. ^ Prentiss, Phoebe G.; W, AV; E, HA (1959-02-28). "Hydropenia in cat and dog. Ability of the cat to meet its water requirements solely from a diet of fish or meat". Am J Physiol 196 (3): 625–632. ISSN 0002-9513. PMID 13627237. 
  85. ^ (Committee on Animal Nutrition 1986, p. 29)
  86. ^ Wolf, A. V.; P, PG; D, LG; S, RJ (1959-02-28). "Potability of sea water with special reference to the cat". Am J Physiol 196 (3): 633–641. ISSN 0002-9513. PMID 13627238. 
  87. ^ Morris JG, Rogers QR, JG (1 December 1978). "Arginine: an essential amino acid for the cat" (Free full text). J. Nutr. 108 (12): 1944–53. ISSN 0022-3166. PMID 722344. 
  88. ^ a b Zoran DL, DL (December 2002). "The carnivore connection to nutrition in cats" (PDF). J. Am. Vet. Med. Assoc. 221 (11): 1559–67. doi:10.2460/javma.2002.221.1559. ISSN 0003-1488. PMID 12479324. 
  89. ^ Gray CM, Sellon RK, Freeman LM, CM (December 2004). "Nutritional adequacy of two vegan diets for cats". J. Am. Vet. Med. Assoc. 225 (11): 1670–5. doi:10.2460/javma.2004.225.1670. ISSN 0003-1488. PMID 15626215. 
  90. ^ a b Zaghini G, Biagi G, G (August 2005). "Nutritional peculiarities and diet palatability in the cat". Vet. Res. Commun. 29 Suppl 2: 39–44. doi:10.1007/s11259-005-0009-1. ISSN 0165-7380. PMID 16244923. 
  91. ^ Some nutritional aspects of ageing in dogs and cats By E. J. Taylor, C. Adams and R. Neville, Proceedings of the Nutrition Society (1995), 54, 645-656
  92. ^ "Me-wow! Texas woman says cat is 30 years old Although she can't hear or see very well, Caterack the cat is still purring". Retrieved 2009-09-30. 
  93. ^ "Cat care: Spay-neuter". American Society for the Prevention of Cruelity to Animals. 2009. Retrieved 2009-09-06. 
  94. ^ Levy JK, Gale DW, Gale LA, JK (January 2003). "Evaluation of the effect of a long-term trap-neuter-return and adoption program on a free-roaming cat population" (PDF). J. Am. Vet. Med. Assoc. 222 (1): 42–6. doi:10.2460/javma.2003.222.42. ISSN 0003-1488. PMID 12523478. 
  95. ^ a b c d e f "Toxic to Cats". Vetinfo4Cats. Retrieved 2007-01-18. 
  96. ^ Williams RT, R. T. (1 February 1978). "Species variations in the pathways of drug metabolism" (Free full text). Environ. Health Perspect. 22: 133–8. doi:10.2307/3428562. ISSN 00916765. PMID 417918. 
  97. ^ Rowland J, J (February 1987). "Incidence of ethylene glycol intoxication in dogs and cats seen at Colorado State University Veterinary Teaching Hospital". Vet Hum Toxicol 29 (1): 41–4. ISSN 0145-6296. PMID 3824875. 
  98. ^ Potera C, C (December 2007). "Chemical exposures: cats as sentinel species". Environ. Health Perspect. 115 (12): A580. ISSN 0091-6765. PMID 18087575. 
  99. ^ Allen AL, AL (1 June 2003). "The diagnosis of acetaminophen toxicosis in a cat". Canadian Veterinary Journal 44 (6): 509–10. ISSN 0008-5286. PMID 12839249. 
  100. ^ Villar D, Buck WB, Gonzalez JM, D (June 1998). "Ibuprofen, aspirin and acetaminophen toxicosis and treatment in Dogs and Cats". Vet Hum Toxicol 40 (3): 156–62. ISSN 0145-6296. PMID 9610496. 
  101. ^ Camille DeClementi; Bailey, Keith L.; Goldstein, Spencer C.; Orser, Michael Scott (December 2004). "Suspected toxicosis after topical administration of minoxidil in 2 cats". Journal of Veterinary Emergency and Critical Care 14 (4): 287–292. doi:10.1111/j.1476-4431.2004.04014.x. 
  102. ^ Bischoff K, Guale F, K (1 April 1998). "Australian tea tree (Melaleuca alternifolia) oil poisoning in three purebred cats". J. Vet. Diagn. Invest. 10 (2): 208–10. ISSN 1040-6387. PMID 9576358. 
  103. ^ Rousseaux CG, Smith RA, Nicholson S, CG (August 1986). "Acute Pinesol toxicity in a domestic cat". Vet Hum Toxicol 28 (4): 316–7. ISSN 0145-6296. PMID 3750813. 
  104. ^ "Antifreeze Warning". The Cat Fanciers' Association, Inc.. Retrieved 2007-05-15. 
  105. ^ "Death by Chocolate? Methylxanthine Toxicosis" (PDF). Veterinary Technician. March 2001. Retrieved 25 August 2009. 
  106. ^ "Plants and Your Cat". The Cat Fanciers' Association, Inc.. Retrieved 2007-05-15. 
  107. ^ Langston, Cathy E (2002-01-01). "Acute renal failure caused by lily ingestion in six cats". Journal of the American Veterinary Medical Association 220 (1): 49–52, 36. doi:10.2460/javma.2002.220.49. ISSN 0003-1488. PMID 12680447. 
  108. ^ Germain, E.; Benhamou, S.; Poulle, M.-L. (2008). "Spatio-temporal sharing between the European wildcat, the domestic cat and their hybrids". Journal of Zoology 276 (2): 195–203. doi:10.1111/j.1469-7998.2008.00479.x. 
  109. ^ a b c Barratt, David G. (1 June 1997). "Home Range Size, Habitat Utilisation and Movement Patterns of Suburban and Farm Cats Felis catus". Ecography 20 (3): 271–280. doi:10.1111/j.1600-0587.1997.tb00371.x. ISSN 09067590. 
  110. ^ Randall, Walter; Johnson, RF; Randall, S; Cunningham, JT (December 1985). "Circadian rhythms in food intake and activity in domestic cats.". Behavioral Neuroscience 99 (6): 1162–1175. doi:10.1037/0735-7044.99.6.1162. ISSN 0735-7044. PMID 3843546. 
  111. ^ Jouvet, Michel (1979). "What does a cat dream about?". Trends in Neurosciences 2: 280-282. doi:10.1016/0166-2236(79)90110-3. ISSN 0166-2236. Retrieved 2010-02-11. 
  112. ^ Pontier, Dominique; Eugenia Natoli (1996). "Male reproductive success in the domestic cat (Felis catus L.): A case history". Behavioural Processes 37 (1): 85–88. doi:10.1016/0376-6357(95)00070-4. ISSN 0376-6357. 
  113. ^ a b Crowell-Davis, SL; Curtis, TM; Knowles, RJ (February 2004). "Social organization in the cat: a modern understanding" (PDF). Journal of Feline Medicine and Surgery 6 (1): 19–28. doi:10.1016/j.jfms.2003.09.013. ISSN 1098-612X. PMID 15123163. Retrieved 2008-05-21. 
  114. ^ Baron, Alan; Stewart, .; Warren, J. M. (1 January 1957). "Patterns of Social Interaction in Cats (Felis domestica)". Behaviour 11 (1): 56–66. doi:10.1163/156853956X00084. ISSN 00057959. Retrieved 2009-08-20. 
  115. ^ a b c d e Bradshaw JW, Goodwin D, Legrand-Defrétin V, Nott HM, JW (July 1996). "Food selection by the domestic cat, an obligate carnivore". Comp. Biochem. Physiol. A Physiol. 114 (3): 205–9. doi:10.1016/0300-9629(95)02133-7. ISSN 1096-4940. PMID 8759144. 
  116. ^ (Jensen 2009, p. 208)
  117. ^ a b "Cat Guide: Body Lanugage". Animal Planet. Retrieved 26 August 2009. 
  118. ^ a b Cafazzo S, Natoli E, S (January 2009). "The social function of tail up in the domestic cat (Felis silvestris catus)". Behav. Processes 80 (1): 60–6. doi:10.1016/j.beproc.2008.09.008. ISSN 0376-6357. PMID 18930121. 
  119. ^ E. Levine; Perry, P; Scarlett, J; Houpt, K (2005). "Intercat aggression in households following the introduction of a new cat" (PDF). Applied Animal Behaviour Science 90 (90): 325–336. doi:10.1016/j.applanim.2004.07.006. Retrieved 2009-04-08. 
  120. ^ Cat Guide: Adolescence and Sexual Maturity Animal Planet
  121. ^ McComb K, Taylor AM, Wilson C, Charlton BD (July 2009). "The cry embedded within the purr". Curr. Biol. 19 (13): R507–8. doi:10.1016/j.cub.2009.05.033. PMID 19602409. 
  122. ^ a b "When Hair Balls Aren't Harmless.". Cornell Feline Health Center. 2006. Retrieved 25 August 2009. 
  123. ^ Boshel, S; Barbazetto, I; Meyer, CH; Peter, S; Stur, M (1982). "Filiform Papillae of Cat Tongue". Cells Tissues Organs 114 (2): 97–105. doi:10.1159/000145583. ISSN 1422-6421. PMID 17728549. Retrieved 2009-08-25. 
  124. ^ a b Lindell, Ellen M. (1997-12). "Intercat aggression: a retrospective study examining types of aggression, sexes of fighting pairs, and effectiveness of treatment". Applied Animal Behaviour Science 55 (1-2): 153–162. doi:10.1016/S0168-1591(97)00032-4. ISSN 0168-1591. Retrieved 2009-08-20. 
  125. ^ Akihiro Yamane, Teruo Doi and Yuiti Ono "Mating behaviors, courtship rank and mating success of male feral cat (Felis catus)" Journal of Ethology, Volume 14, Number 1, p35-44 (1996) doi:10.1007/BF02350090
  126. ^ Aggression Between Family Cats The Humane Society of the United States 2002
  127. ^ Pedersen NC, Yamamoto JK, Ishida T, Hansen H, NC (May 1989). "Feline immunodeficiency virus infection". Vet. Immunol. Immunopathol. 21 (1): 111–29. doi:10.1016/0165-2427(89)90134-7. ISSN 0165-2427. PMID 2549690. 
  128. ^ a b c Woods M, McDonald RA, Harris S. (2003). "Predation of wildlife by domestic cats Felis catus in Great Britain". Mammal Review 23 (2): 174–188. doi:10.1046/j.1365-2907.2003.00017 (inactive 2009-09-06). 
  129. ^ a b (Turner & Bateson 2000, p. 153)
  130. ^ "Why Do Cats Like High Places?". Drs. Foster & Smith, Inc.. Dr. Holly Nash, DVM, MS. Archived from the original on 2008-01-02. 
  131. ^ "Falling Cats". Archived from the original on 2005-10-26. Retrieved 2005-10-24. 
  132. ^ Huy D. Nguyen. "How does a Cat always land on its feet?". Georgia Institute of Technology, School of Medical Engineering. Retrieved 2007-05-15. 
  133. ^ Leyhausen, Paul (1978). Cat Behavior: The Predatory and Social Behavior of Domestic and Wild Cats. New York: Garland STPM Press. ISBN 978-0824070175. 
  134. ^ Desmond, Morris (1986). Catwatching Why Cats Purr and Everything Else You Ever Wanted to Know. Crown Pub. 
  135. ^ Kienzle E, E (December 1994). "Blood sugar levels and renal sugar excretion after the intake of high carbohydrate diets in cats" (PDF). J. Nutr. 124 (12 Suppl): 2563S–2567S. ISSN 0022-3166. PMID 7996238. 
  136. ^ Bradshaw, John W. S. (1997-04). "Factors affecting pica in the domestic cat". Applied Animal Behaviour Science 52 (3-4): 373–379. doi:10.1016/S0168-1591(96)01136-7. ISSN 0168-1591. Retrieved 2009-08-20. 
  137. ^ "Pica: The Un-finicky Feline". University of California School of Veterinary Medicine. Retrieved 2009-09-06. 
  138. ^ Poirier FE, Hussey LK, F. E. (1 July 1982). "Nonhuman Primate Learning: The Importance of Learning from an Evolutionary Perspective". Anthropology & Education Quarterly 13 (2): 133–148. doi:10.1525/aeq.1982.13.2.05x1830j. ISSN 01617761. 
  139. ^ Byers, John A.; Bekoff, Marc (1998). Animal play: evolutionary, comparative, and ecological perspectives. Cambridge, UK: Cambridge University Press. p. 135. ISBN 0-521-58656-9. 
  140. ^ Hall, Sarah L. (2002-11-01). "Object play in adult domestic cats: the roles of habituation and disinhibition". Applied Animal Behaviour Science 79 (3): 263–271. doi:10.1016/S0168-1591(02)00153-3. ISSN 0168-1591. Retrieved 2009-08-20. 
  141. ^ Hall, Sarah L. (1998-06). "The influence of hunger on object play by adult domestic cats". Applied Animal Behaviour Science 58 (1-2): 143–150. doi:10.1016/S0168-1591(97)00136-6. ISSN 0168-1591. Retrieved 2009-08-20. 
  142. ^ MacPhail, ST (2002-11). "Gastrointestinal obstruction". Clinical Techniques in Small Animal Practice 17 (4): 178–183. doi:10.1053/svms.2002.36606. ISSN 1096-2867. PMID 12324686. PMC 2391006. Retrieved 2009-08-20. 
  143. ^ "Fat indoor cats need exercise". Pocono Record. December 10, 2006. 
  144. ^ Eye expert challenges laser pen 'danger' BBC News November 24, 1997
  145. ^ a b c d e "Prolific Cats: The Estrous Cycle" (PDF). Veterinary Learning Systems. Retrieved 19 June 2009. 
  146. ^ Aronson LR, Cooper ML, LR (January 1967). "Penile spines of the domestic cat: their endocrine-behavior relations" (PDF). Anat. Rec. 157 (1): 71–8. doi:10.1002/ar.1091570111. ISSN 0003-276X. PMID 6030760. 
  147. ^ Wildt DE, Seager SW, Chakraborty PK, DE (October 1980). "Effect of copulatory stimuli on incidence of ovulation and on serum luteinizing hormone in the cat". Endocrinology 107 (4): 1212–7. doi:10.1210/endo-107-4-1212. ISSN 0013-7227. PMID 7190893. 
  148. ^ Tsutsui T, Stabenfeldt GH, T (1993). "Biology of ovarian cycles, pregnancy and pseudopregnancy in the domestic cat". J. Reprod. Fertil. Suppl. 47: 29–35. ISSN 0449-3087. PMID 8229938. 
  149. ^ Behrend, Katrin; Wegler, Monika; translated from German by Elizabeth D. Crawford. (1991). The Complete Book of Cat Care: How to Raise a Happy and Healthy Cat. Hauppauge, NY: Barron's Educational Series, Inc.. p. 28. ISBN 0-8120-4613-7. 
  150. ^ Olson PN, Kustritz MV, Johnston SD, PN (2001). "Early-age neutering of dogs and cats in the United States (a review)". J. Reprod. Fertil. Suppl. 57: 223–32. ISSN 0449-3087. PMID 11787153. 
  151. ^ Margaret V. Root Kustritz "Determining the optimal age for gonadectomy of dogs and cats." J Am Vet Med Assoc. 2007 Dec 1;231(11):1665-75
  152. ^ Chu, K; Anderson, WM; Rieser, MY (April 2009). "Population characteristics and neuter status of cats living in households in the United States". Journal of the American Veterinary Medical Association 234 (8): 1023–1030. doi:10.2460/javma.234.8.1023. ISSN 0003-1488. PMID 19366332. Retrieved 2009-08-21. 
  153. ^ Say, Ludovic (2002-02-01). "Spatio-temporal variation in cat population density in a sub-Antarctic environment". Polar Biology 25 (2): 90–95. doi:10.1007/s003000100316. 
  154. ^ Frenot, Yves; Chown, Steven L.; Whinam, Jennie; Selkirk, Patricia M.; Convey, Peter; Skotnicki, Mary; Bergstrom, Dana M. (2005). "Biological Invasions in the Antarctic: Extent, Impacts and Implications". Biological Reviews 80 (01): 45–72. doi:10.1017/S1464793104006542. Retrieved 2009-08-28. 
  155. ^ "Global Invasive Species Database: Ecology of Felis catus". Invasive Species Specialist Group. 2006. Retrieved 31 August 2009. 
  156. ^ a b Nogales M, Martin A, Tershy BR, Donlan CJ, Veitch D, Uerta N, Wood B, Alonso J. (2004). "A Review of Feral Cat Eradication on Islands.". Conservation Biology 18 (2): 310. doi:10.1111/j.1523-1739.2004.00442.x. 
  157. ^ "100 of the World's Worst Invasive Alien Species: a Selection from the Global Invasive Species Database" (PDF). The Invasive Species Specialist Group. 2000. Archived from the original on 2007-09-27. Retrieved 31 August 2009. 
  158. ^ May, R. (1988). "Control of feline delinquency". Nature 332: 392–393. 
  159. ^ Liberg, O. (1982). "Food habits and prey impact by feral and house-based domestic cats in a rural area in southern Sweden". Journal of Mammalogy 65: 424–432. 
  160. ^ Robertson ID, ID (August 1998). "Survey of predation by domestic cats". Aust. Vet. J. 76 (8): 551–4. doi:10.1111/j.1751-0813.1998.tb10214.x. ISSN 0005-0423. PMID 9741724. 
  161. ^ Beckerman AP, Boots M, Gaston KJ. (2007). "Urban bird declines and the fear of cats" (PDF). Animal Conservation 10 (3): 320–325. doi:10.1111/j.1469-1795.2007.00115.x. 
  162. ^ Courchamp F, Chapuis JL, Pascal M, F (August 2003). "Mammal invaders on islands: impact, control and control impact". Biol Rev Camb Philos Soc 78 (3): 347–83. doi:10.1017/S1464793102006061. ISSN 1464-7931. PMID 14558589. 
  163. ^ Rayner MJ, Hauber ME, Imber MJ, Stamp RK, Clout MN, MJ (December 2007). "Spatial heterogeneity of mesopredator release within an oceanic island system" (Free full text). Proc. Natl. Acad. Sci. U.S.A. 104 (52): 20862–5. doi:10.1073/pnas.0707414105. ISSN 0027-8424. PMID 18083843. PMC 2409232. 
  164. ^ Chris R. Dickman (1996). "Overview of the Impacts of Feral Cats on Australian Native Fauna" (PDF). Australian Department of the Environment, Water, Heritage and the Arts. Retrieved 28 August 2009. 
  165. ^ James H, Acharya AB, Taylor JA, Freak MJ, H (June 2002). "A case of bitten Bettongs". J Forensic Odontostomatol 20 (1): 10–2. ISSN 0258-414X. PMID 12085522. 
  166. ^ Glen AS, Dickman CR, AS (August 2005). "Complex interactions among mammalian carnivores in Australia, and their implications for wildlife management". Biol Rev Camb Philos Soc 80 (3): 387–401. doi:10.1017/S1464793105006718. ISSN 1464-7931. PMID 16094805. 
  167. ^ Chucher, P.B.; Lawton, J.H. (1987). "Predation by domestic cats in an English village.". Journal of Zoology, London 212: 439–455. 
  168. ^ a b Mead, C.J. (1982). "Ringed birds killed by cats". Mammal Review 12(4): 183–186. 
  169. ^ Fiztgerald, M.B.; Turner, D.C.. Hunting behaviour of domestic cats and their impact on prey populations. In: The Domestic Cat: The biology of its behaviour, 2nd ed. Turner, D.C. and Bateson, P. (eds.),. Cambridge: Cambridge University Press. p. 151–175. 
  170. ^ Courchamp, F.; Langlais, M. and Sugihara, G. (1999). "Cats protecting birds: modelling the mesopredator release effect". Mammal Review 12(4): 183–186. 
  171. ^ Stattersfield, A.J.; Crosby, M.J., Long, A.J. and Wege, D.C. (1998). Endemic Bird Areas of the World: Priorities for Biodiversity Conservation.. Cambridge.: BirdLife Conservation Series No.7. , Burlington Press. 
  172. ^ Williams, A.J. (1984). Status and Conservation of Seabirds at some Islands in the African Sector of the Southern Ocean. In: Status and Conservation of the World’s Seabirds, ed Croxall, J.P., Evans, P.G.H. & Schreiber, R.W. International Council for Bird Preservation.. Cambridge.. p. 627–635. 
  173. ^ Falla, R.A. (1955). Zealand Bird Life past and present. Stiles.. 
  174. ^ Galbreath, R.; Brown, D. (2004). "The tale of the lighthouse-keeper’s cat: discovery and extinction of the Stephens Island wren (Traversia lyalli).". Notornis 51: 193-200. 
  175. ^ a b Dowding, J.E.; Murphy, E.C. (2001). "The Impact of Predation be Introduced Mammals on Endemic Shorebirds in New Zealand: A Conservation Perspective.". Biological Conservation 99: 47–64. 
  176. ^ Whiting, M.F.; Bradler, S. and Maxwell, T. (2003). "Loss and Recovery of Wings in Stick Insects". Nature 421: 264–267. 
  177. ^ WCMC, . (1992). World Conservation Monitoring Centre.. Chapman and Hall.. 
  178. ^ Steadman, D.W.; Martin, P.S. (2003). "The Late Quaternary Extinction and Future Resurrection of Birds on Pacific Islands". Earth Science Reviews 61: 133–147. 
  179. ^ (Turner & Bateson 2000, p. 4)
  180. ^ a b "Market research statistics - U.S. pet ownership". American Veterinary Medical Association. Retrieved 27 August 2009. 
  181. ^ Abby Ellin. "More Men Are Unabashedly Embracing Their Love of Cats". New York Times. Retrieved 30 August 2009. 
  182. ^ a b Jeffrey M. Jones. "Companionship and Love of Animals Drive Pet Ownership". Gallup, Inc. Retrieved 30 August 2009. 
  183. ^ ASPCA Complete Guide to Cats by James R. Richards,, DVM
  184. ^ "What is that they’re wearing?" (Pdf). Humane Society of the United States. Retrieved 22 October 2009. 
  185. ^ "EU proposes cat and dog fur ban". BBC News. Retrieved 22 October 2009. 
  186. ^ "EU Announces Strict Ban on Dog and Cat Fur Imports and Exports". Humane Society International. Retrieved 22 October 2009. 
  187. ^ "Switzerland finds a way to skin a cat for the fur trade and high fashion". The Independent. Retrieved 23 October 2009. 
  188. ^ "China protesters: Stop 'cooking cats alive'." December 18, 2008.
  189. ^ "How many dogs and cats are eaten in Asia?". September 2003.
  190. ^
  191. ^ (Mason 1984, p. 224)
  192. ^ "Moggie". Retrieved 2007-05-15. 
  193. ^ "Torties, Calicos and Tricolor cats". Retrieved 2005-10-24. 
  194. ^ a b Kravetz JD, Federman DG, JD (September 2002). "Cat-associated zoonoses" (Free full text). Arch. Intern. Med. 162 (17): 1945–52. doi:10.1001/archinte.162.17.1945 (inactive 2009-10-23). ISSN 0003-9926. PMID 12230416. 
  195. ^ Talan DA, Citron DM, Abrahamian FM, Moran GJ, Goldstein EJ, DA (January 1999). "Bacteriologic analysis of infected dog and cat bites. Emergency Medicine Animal Bite Infection Study Group". N. Engl. J. Med. 340 (2): 85–92. doi:10.1056/NEJM199901143400202. ISSN 0028-4793. PMID 9887159. 
  196. ^ Torda A, A (August 2001). "Toxoplasmosis. Are cats really the source?". Aust Fam Physician 30 (8): 743–7. ISSN 0300-8495. PMID 11681144. 
  197. ^ Svobodová V, Knotek Z, Svoboda M, V (December 1998). "Prevalence of IgG and IgM antibodies specific to Toxoplasma gondii in cats". Vet. Parasitol. 80 (2): 173–6. doi:10.1016/S0304-4017(98)00201-5. ISSN 0304-4017. PMID 9870370. 
  198. ^ Meireles LR, Galisteo AJ, Pompeu E, Andrade HF, LR (August 2004). "Toxoplasma gondii spreading in an urban area evaluated by seroprevalence in free-living cats and dogs" (Free full text). Trop. Med. Int. Health 9 (8): 876–81. doi:10.1111/j.1365-3156.2004.01280.x. ISSN 1360-2276. PMID 15303992. 
  199. ^ De Craeye S, Francart A, Chabauty J, S (October 2008). "Prevalence of Toxoplasma gondii infection in Belgian house cats". Vet. Parasitol. 157 (1-2): 128–32. doi:10.1016/j.vetpar.2008.07.001. ISSN 0304-4017. PMID 18707811. 
  200. ^ Erwin EA, Woodfolk JA, Custis N, Platts-Mills TA, EA (August 2003). "Animal danders". Immunol Allergy Clin North Am 23 (3): 469–81. doi:10.1016/S0889-8561(03)00004-3. ISSN 0889-8561. PMID 14524386. 
  201. ^ "Dealing with cat allergies" (PDF). Archived from the original on 2007-06-04. 
  202. ^ Simpson A, Custovic A, A (February 2003). "Early pet exposure: friend or foe?". Curr Opin Allergy Clin Immunol 3 (1): 7–14. doi:10.1097/01.all.0000053261.39029.7a (inactive 2009-09-06). ISSN 1528-4050. PMID 12582308. 
  203. ^ Simpson A, Custovic A, A (May 2005). "Pets and the development of allergic sensitization". Curr Allergy Asthma Rep 5 (3): 212–20. doi:10.1007/s11882-005-0040-x. ISSN 1529-7322. PMID 15842959. 
  204. ^ Avner, D B; Perzanowski, MS; Platts-Mills, TA; Woodfolk, JA (1997-09). "Evaluation of different techniques for washing cats: quantitation of allergen removed from the cat and the effect on airborne Fel d 1". The Journal of Allergy and Clinical Immunology 100 (3): 307–312. doi:10.1016/S0091-6749(97)70242-2. ISSN 0091-6749. PMID 9314341. 
  205. ^ Miller, H (February 2005). "Cat and mouse in regulating genetic 'enhancement'". Nature Biotechnology 23 (2): 171–172. doi:10.1038/nbt0205-171. ISSN 1087-0156. PMID 15696141. Retrieved 2009-08-21. 
  206. ^ Allen K, Blascovich J, Mendes WB, K (September 2002). "Cardiovascular reactivity and the presence of pets, friends, and spouses: the truth about cats and dogs" (Free full text). Psychosom Med 64 (5): 727–39. doi:10.1097/01.PSY.0000024236.11538.41. ISSN 0033-3174. PMID 12271103. 
  207. ^ (Fogle 1981, pp. 23–56)
  208. ^ Landsberg GM, GM (March 1991). "Feline scratching and destruction and the effects of declawing". Vet. Clin. North Am. Small Anim. Pract. 21 (2): 265–79. ISSN 0195-5616. PMID 2053250. 
  209. ^ "Scratching or clawing in the house". Retrieved 2005-08-14. 
  210. ^ a b Swiderski J, J (November 2002). "Onychectomy and its alternatives in the feline patient". Clin Tech Small Anim Pract 17 (4): 158–61. doi:10.1053/svms.2002.36604. ISSN 1096-2867. PMID 12587280. 
  211. ^ Welfare Implications of Declawing of Domestic Cats American Veterinary Medical Association 09 April 2009
  212. ^ Patronek GJ, GJ (October 2001). "Assessment of claims of short- and long-term complications associated with onychectomy in cats" (Free full text). J. Am. Vet. Med. Assoc. 219 (7): 932–7. doi:10.2460/javma.2001.219.932. ISSN 0003-1488. PMID 11601788. 
  213. ^ Paw Project Acknowledgements
  214. ^ Hornfeldt, CS; Westfall (1996). "Suspected bentonite toxicosis in a cat from ingestion of clay cat litter.". Veterinary and human toxicology 38 (5): 365–6. PMID 8888544. 
  215. ^ "Parasite in cats killing sea otters". NOAA magazine (National Oceanic and Atmospheric Administration). 2003-01-21. Retrieved 2007-11-24. 
  216. ^ Brown-Martin, Darcy (March 2007). "Monterey Bay’s sea otter sleuth". Retrieved 2008-11-22. 
  217. ^ "A Step-By-Step Guide on How To Train Your Cat to Use the Human Toilet". The Toilet Trained Cat. Retrieved 2009-02-15. 
  218. ^ a b Irene Rochlitz (2007). The Welfare of Cats (Animal Welfare). Berlin: Springer. pp. 141–175. ISBN 1-4020-6143-9. 
  219. ^ a b c What is the difference between a stray cat and a feral cat? Humane Society of the United States
  220. ^ Torre Argentina cat shelter. Retrieved 17 June 2009.
  221. ^ Rowan, Andrew N.; Deborah J. Salem (2003-11). "4" (PDF). The State of the Animals II: 2003. Humane Society Press. ISBN 0965894274. 
  222. ^ Muir, Hazel (2004-04-08). "Ancient remains could be oldest pet cat". New Scientist. Retrieved 2007-11-23. 
  223. ^ Walton, Marsha (April 9, 2004). "Ancient burial looks like human and pet cat". CNN. Retrieved 2007-11-23. 
  224. ^ (Mason 1984, p. 220)
  225. ^ a b (Mason 1984, p. 223)
  226. ^ Geyer, Georgie Anne (2004). When Cats Reigned Like Kings: On the Trail of the Sacred Cats. Kansis City: Andrews McMeel Pub.. ISBN 0740746979. 
  227. ^ Minou Reeves (2000). Muhammad in Europe. New York University (NYU) Press. p. 52. ISBN 0814775330. 
  228. ^ "Cat Myths, Misinformation and Untruths". Retrieved 2008-11-22. 
  229. ^ The ASPCA Warns About High-Rise Falls by Cats.

Further reading

  • Case, Linda P. (2003). The cat: its behavior, nutrition, & health. Ames, Iowa: Iowa State Press. ISBN 0-8138-0331-4. 
  • Committee on Animal Nutrition (1986). Nutrient Requirements of Cats (2nd ed.). National Academy Press. ISBN 0309074835. . 
  • Fogle (1981).^ By on October 22, 2009 at 5:36 am the good happens blog… .
    • Stay Dougie » Mos Def In Your Neighborhood Boneyard With Stussy 11 January 2010 15:14 UTC [Source type: FILTERED WITH BAYES]

    ^ By on December 2, 2009 at 3:27 pm www food saver… .
    • Stay Dougie » Mos Def In Your Neighborhood Boneyard With Stussy 11 January 2010 15:14 UTC [Source type: FILTERED WITH BAYES]

    ^ By on October 29, 2009 at 9:21 am food 4 less stores angeles… .
    • Stay Dougie » Mos Def In Your Neighborhood Boneyard With Stussy 11 January 2010 15:14 UTC [Source type: FILTERED WITH BAYES]

    Interrelations Between People and Pets. Charles C Thomas Pub Ltd. ISBN 0398041695. 
  • Jensen, Per (2009). The Ethology of Domestic Animals (Modular Text). Wallingford: CABI. ISBN 1-84593-536-5. 
  • Kahn, Cynthia M.; Line, Scott (2007). The Merck/Merial Manual for Pet Health (1 ed.). Merck. ISBN 0911910999. 
  • Mason, I.L. (1984). Evolution of Domesticated Animals. Prentice Hall Press. ISBN 0582460468. 
  • Rochlitz, Irene (2007). The Welfare of Cats (1 ed.). Springer. ISBN 1402061439. 
  • Sunquist, Melvin E; Sunquist, Fiona (2002). Wild cats of the world. University of Chicago Press. ISBN 0226779998. 
  • Turner, D.C.; Bateson, P. (2000). The Domestic Cat: The Biology of its Behaviour (2 ed.). Cambridge University Press. ISBN 0521636485. 
  • Walker, Warren F. (1982). Study of the Cat with Reference to Human Beings (4th Revised ed.). Thomson Learning. ISBN 0030579147. 

External links

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Up to date as of January 14, 2010

From Wikiquote

This entry is about the animal. For the musical, see Cats (musical).
In ancient times, cats were worshipped as gods. They have never forgotten this. ~ Terry Pratchett
The cat, also called the domestic cat or house cat, is a small feline carnivorous mammal of the subspecies Felis silvestris catus. It has been living in close association with humans for between 3,500 and 8,000 years.



  • I think all cats are wild. They only act tame if there's a saucer of milk in it for them.
  • Some see "God" as "dog" spelled backwards. I see "God" as "cat" spelled with a vivid imagination.
  • Lat take a cat, and fostre hym wel with milk
    And tendre flessh, and make his couche of silk,
    And lat hym seen a mous go by the wal,
    Anon he weyveth milk and flessh and al,
    And every deyntee that is in that hous,
    Swich appetit hath he to ete a mous.
  • Anyone who considers protocol unimportant has never dealt with a cat.
  • Perhaps God made cats so that man might have the pleasure of fondling the tiger...
The kitten has a luxurious, Bohemian, unpuritanical nature. ~ Robertson Davies
  • The kitten has a luxurious, Bohemian, unpuritanical nature. It eats six meals a day, plays furiously with a toy mouse and a piece of rope, and suddenly falls into a deep sleep whenever the fit takes it. It never feels the necessity to do anything to justify its existence; it does not want to be a Good Citizen; it has never heard of Service. It knows that it is beautiful and delightful, and it considers that a sufficient contribution to the general good. And in return for its beauty and charm it expects fish, meat, and vegetables, a comfortable bed, a chair by the grate fire, and endless petting.
The Naming of cats is a difficult matter; It isn't just one of your holiday games. ~ T. S. Eliot
  • The Naming of cats is a difficult matter;
    It isn't just one of your holiday games.
    You may think at first, I'm as mad as a hatter
    When I tell you a cat must have three different names.
  • Nothing's more playful than a young cat, nor more grave than an old one.
  • Cat, I'm a kitty-cat, and I dance, dance, dance, and I dance, dance, dance.
    • Steve Ibsen, "The Kitty Cat Dance"
  • A cat who turns her nose up at bread does not deserve meat.
  • It is said that in Ulthar, which lies beyond the river Skai, no man may kill a cat; and this I can verily believe as I gaze upon him who sitteth purring before the fire. For the cat is cryptic, and close to strange things which men cannot see. He is the soul of antique Aegyptus, and bearer of tales from forgotten cities in Meroe and Ophir. He is the kin of the jungle’s lords, and heir to the secrets of hoary and sinister Africa. The Sphinx is his cousin, and he speaks her language; but he is more ancient than the Sphinx, and remembers that which she hath forgotten.
  • Don't want a cat
    Scratching its claws all over my
    Giving no love and getting fat
    Oh, you can get lonely
    And a cat's no help with that.

In praise of cats

A thing of beauty, strength, and grace lies behind that whiskered face - Anonymous
  • For I will consider my Cat Jeoffry.
    For he is the servant of the Living God duly and daily serving him.
    For at the first glance of the glory of God in the East he worships in his way.
    For this is done by wreathing his body seven times round with elegant quickness.
    • Christopher Smart (1722-1771), "For I will consider my Cat Jeoffry" from "Jubilate Agno" (1758-1763)
  • When there was room on the ledge outside of the pots and boxes for a cat, the cat was there — in sunny weather — stretched at full length, asleep and blissful, with her furry belly to the sun and a paw curved over her nose. Then that house was complete, and its contentment and peace were made manifest to the world by this symbol, whose testimony is infallible. A home without a cat—and a well-fed, well-petted, and properly revered cat—may be a perfect home, perhaps, but how can it prove title?
In the eyes of a cat, all things belong to cats.

Cats and humans

  • Authors like cats because they are such quiet, lovable, wise creatures, and cats like authors for the same reasons.
  • The cat is a wild animal that inhabits the homes of humans.
    • Konrad Lorenz, Man Meets Dog


Dogs have Masters. Cats have staff.
  • Our character is what God and cats know of us.
Cats, no less liquid than their shadows, offer no angles to the wind...
  • When the rat laughs at the cat, there is a hole. The rat has not power to call the cat to account. The rat does not go to sleep in the cat's bed.
    • African Proverb
  • In the eyes of a cat, all things belong to cats.
  • No matter how much cats fight, there always seem to be plenty of kittens.
  • Siamese Cats have a way of staring at you. Those who have walked in on the Queen cleaning her teeth will know the expression.
  • No matter if it is a white cat or a black cat; as long as it can catch mice, it is a good cat.

In praise of cats

  • A meow massages the heart.
    • Stuart McMillan
  • Do not meddle in the affairs of cats, for they are subtle and will pee on your computer.
    • Bruce Graham
    • echoing Tolkien's "Do not meddle in the affairs of wizards, for they are subtle and quick to anger."
    • Cats will purr at content or when they are feeling frisky.

Cats and humans

  • Dogs have Masters. Cats have staff.
    • Anonymous
  • Cats seem to go on the principle that it never does any harm to ask for what you want.
    • Joseph Wood Krutch
  • I gave my cat a bath the other day... They love it. He sat there, he enjoyed it, it was fun for me. The fur would stick to my tongue, but other than that...
  • You can tell your cat anything and he'll still love you. If you lose your job or your best friend, your cat will think no less of you.
    • Helen Powers

Cat elegance

  • Cats, no less liquid than their shadows, offer no angles to the wind. They slip, diminished, neat, through loopholes less than themselves.

Cat arrogance

  • Cats are smarter than dogs. You can't get eight cats to pull a sled through snow.
    • Jeff Valdez
  • In ancient times, cats were worshiped as gods. They have never forgotten this.
  • A cat knows exactly what you are, and treats you accordingly.
    • author unknown
Everyone knows cats are on a higher level of existence. These silly humans are just too big-headed to admit their inferiority.
  • Cats understand our feelings. They don't care, but they understand.

Cat intelligence

  • Cats are experiments in consciousness with limited intelligence, humans are experiments in intelligence with limited consciousness (unless they work on themselves).
  • Children are like cats, they can tell when you don't like them. That's when they come over and violate your body space.
  • Everyone knows cats are on a higher level of existence. These silly humans are just too big-headed to admit their inferiority. Just think what a nicer world this would be if it were controlled by cats.
    You wouldn't see cats having waste disposal problems. They're neat.
    They don't have sexual hangups. A cat gets horny, it does something about it.
    They keep reasonable hours. You never see a cat up before noon.
    They know how to relax. Ever heard of a cat with an ulcer?
    What are the chances of a cat starting a nuclear war? Pretty neglible. It's not that they can't, they just know that there are much better things to do with one's time. Like lie in the sun and sleep. Or go exploring the world.
  • I have studied many philosophers and many cats. The wisdom of cats is infinitely superior.

External links

Wikipedia has an article about:
Look up cat in Wiktionary, the free dictionary

Source material

Up to date as of January 22, 2010
(Redirected to The Cats article)

From Wikisource

The Cats
by H. P. Lovecraft
Babels of blocks to the high heavens towering
Flames of futility swirling below;
Poisonous fungi in brick and stone flowering,
Lanterns that shudder and death-lights that glow.
Black monstrous bridges across oily rivers,
Cobwebs of cable to nameless things spun;
Catacomb deeps whose dank chaos delivers
Streams of live foetor that rots in the sun.
Colour and splendour, disease and decaying,
Shrieking and ringing and crawling insane,
Rabbles exotic to stranger-gods praying,
Jumbles of odour that stifle the brain.
Legions of cats from the alleys nocturnal.
Howling and lean in the glare of the moon,
Screaming the future with mouthings infernal,
Yelling the Garden of Pluto's red rune.
Tall towers and pyramids ivy'd and crumbling,
Bats that swoop low in the weed-cumber'd streets;
Bleak Arkham bridges o'er rivers whose rumbling
Joins with no voice as the thick horde retreats.
.Belfries that buckle against the moon totter,
Caverns whose mouths are by mosses effac'd,
And living to answer the wind and the water,
Only the lean cats that howl in the wastes.
^ Sissy- Kitty My Sweet Cat published by myahsan My nice cats Precious the Cat published by moldybean6 This is the only cat that came to live in the house with us after we sold the cow.
  • cat pictures, videos and albums 11 September 2009 10:29 UTC [Source type: General]


Up to date as of January 15, 2010

Definition from Wiktionary, a free dictionary



  1. Plural form of Cat.


Citable sentences

Up to date as of December 12, 2010

Here are sentences from other pages on Lion, which are similar to those in the above article.

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