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The Cryptomonads-haptophytes assemblage is a monophyletic grouping[1] of chromalveolata that are not included in the SAR supergroup.

It includes:

In the past, heterokonts, haptophytes, and cryptomonads have been sometimes grouped together in a group known as chromist.[2] Though the heterokonts are now split out, Cryptophyta and Haptophyta are considered to be closely related[3] (and are sometimes simply referred to as the "Cryptophyta+Haptophyta" group.)[4] A 2009 paper also suggests that the Telonemia and centrohelids may form a clade with the cryptophytes and haptophytes.[5] The picobiliphytes may belong in this group but are too poorly known to be classified with confidence.[6]

The term "Hacrobia" has been proposed for this group.[6] The "Ha-" refers to haptophyta, the "-cr-" refers to cryptomonads, and the "-bia" is a general suffix referring to life.


  1. ^ Sakaguchi M, Takishita K, Matsumoto T, Hashimoto T, Inagaki Y (July 2009). "Tracing back EFL gene evolution in the cryptomonads-haptophytes assemblage: separate origins of EFL genes in haptophytes, photosynthetic cryptomonads, and goniomonads". Gene 441 (1-2): 126–31. doi:10.1016/j.gene.2008.05.010. PMID 18585873.  
  2. ^ Csurös M, Rogozin IB, Koonin EV (May 2008). "Extremely intron-rich genes in the alveolate ancestors inferred with a flexible maximum-likelihood approach". Mol. Biol. Evol. 25 (5): 903–11. doi:10.1093/molbev/msn039. PMID 18296415.  
  3. ^ Rice DW, Palmer JD (2006). "An exceptional horizontal gene transfer in plastids: gene replacement by a distant bacterial paralog and evidence that haptophyte and cryptophyte plastids are sisters". BMC Biol. 4: 31. doi:10.1186/1741-7007-4-31. PMID 16956407. PMC 1570145.  
  4. ^ Reeb VC, Peglar MT, Yoon HS, et al. (May 2009). "Interrelationships of chromalveolates within a broadly sampled tree of photosynthetic protists". Mol. Phylogenet. Evol.. doi:10.1016/j.ympev.2009.04.012. PMID 19398025.  
  5. ^ Burki, F; Inagaki, Y; Bråte, J; Archibald, J.; Keeling, P.; Cavalier-Smith, T; Sakaguchi, M; Hashimoto, T; Horak, A; Kumar, S; Klaveness, D; Jakobsen, K.S; Pawlowski, J; Shalchian-Tabrizi, K (2009). "Large-scale phylogenomic analyses reveal that two enigmatic protist lineages, Telonemia and Centroheliozoa, are related to photosynthetic chromalveolates." (Free full text). Genome Biology and Evolution. doi:doi:10.1093/gbe/evp022.  
  6. ^ a b Okamoto, N.; Chantangsi, C.; Horák, A.; Leander, B.; Keeling, P. (2009). "Molecular Phylogeny and Description of the Novel Katablepharid Roombia truncata gen. Et sp. Nov., and Establishment of the Hacrobia Taxon nov". PloS one 4 (9): e7080. doi:10.1371/journal.pone.0007080. PMID 19759916.   edit

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