Radiolarian: Wikis

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Radiolaria
Fossil range: Cambrian - Recent
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Radiolaria illustration from the Challenger Expedition 1873-76.
Scientific classification
Domain: Eukaryota
Kingdom: Rhizaria
Superphylum: Retaria
Phylum: Radiolaria
Müller 1858 emend.
Classes

Polycystinea
Acantharea
Sticholonchea

Radiolarians (also radiolaria) are amoeboid protozoa that produce intricate mineral skeletons, typically with a central capsule dividing the cell into inner and outer portions, called endoplasm and ectoplasm. They are found as zooplankton throughout the ocean, and their skeletal remains cover large portions of the ocean bottom as radiolarian ooze. Due to their rapid turn-over of species, they represent an important diagnostic fossil found from the Cambrian onwards. Some common radiolarian fossils include Actinomma, Heliosphaera and Hexadoridium.

Contents

Description

Circogonia icosahedra, a species of Radiolaria, shaped like a regular icosahedron

Radiolarians have many needle-like pseudopodia supported by bundles of microtubules, called axopods, which aid in the Radiolarian's buoyancy. The nuclei and most other organelles are in the endoplasm, while the ectoplasm is filled with frothy vacuoles and lipid droplets, keeping them buoyant. Often it also contains symbiotic algae, especially zooxanthellae, which provide most of the cell's energy. Some of this organization is found among the heliozoa, but those lack central capsules and only produce simple scales and spines.

The main class of radiolarians are the Polycystinea, which produce siliceous skeletons. These include the majority of fossils. They also include the Acantharea, which produce skeletons of strontium sulfate. Despite some initial suggestions to the contrary, genetic studies place these two groups close together. They also include the peculiar genus Sticholonche, which lacks an internal skeleton and so is usually considered a heliozoan.

Traditionally the radiolarians have also included the Phaeodarea, which produce siliceous skeletons but differ from the polycystines in several other respects. However, on molecular trees they branch with the Cercozoa, a group including various flagellate and amoeboid protists.
The other radiolarians appear near, but outside, the Cercozoa, so the similarity is due to convergent evolution. The radiolarians and Cercozoa are included within a supergroup called the Rhizaria.

Some radiolarians are known for their resemblance to regular polyhedra, such as with the icosohedron-shaped Circogonia icosahedra pictured to the left.

Fossil record

The earliest known radiolaria date to the very start of the Cambrian period, appearing in the same beds as the first small shelly fauna - they may even be terminal precambrian in age. They differ little from later radiolaria.[1]

Haeckel's radiolarians

German biologist Ernst Haeckel produced exquisite (and perhaps somewhat exaggerated) drawings of radiolaria, helping to popularize these protists among Victorian parlor microscopists alongside foraminifera and diatoms.

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Illustrations from Haeckel's Kunstformen der Natur (1904)

See also

References

  1. ^ Braun, Chen, Waloszek & Maas (2007), "First Early Cambrian Radiolaria", in Vickers-Rich, Patricia; Komarower, Patricia, The Rise and Fall of the Ediacaran Biota, Special publications, 286, London: Geological Society, pp. 143–149, doi:10.1144/SP286.10, ISBN 9781862392335, OCLC 156823511 191881597  
  • Zettler, Linda A. (1997). "Phylogenetic relationships between the Acantharea and the Polycystinea: A molecular perspective on Haeckel's Radiolaria". Proceedings of the National Academy of Sciences 94: 11411–11416. doi:10.1073/pnas.94.21.11411. PMID 9326623.  
  • P. Lopez-Garcia et al. (2002). "Toward the Monophyly of Haeckel's Radiolaria: 18S rRNA Environmental Data Support the Sisterhood of Polycystinea and Acantharea". Molecular Biology and Evolution 19 (1): 118–121.  
  • Sina M. Adl et al. (2005). "The New Higher Level Classification of Eukaryotes with Emphasis on the Taxonomy of Protists". Journal of Eukaryotic Microbiology 52 (5): 399–451. doi:10.1111/j.1550-7408.2005.00053.x.  
  • Haeckel, Ernst (2005). Art Forms from the Ocean: The Radiolarian Atlas of 1862. Munich; London: Prestel Verlag. ISBN 3-7913-3327-5.  

External links


Simple English

File:Haeckel
Radiolarian shells are extremely varied. This is just one family, the Cyrtoidea.

Radiolarians (Radiolaria) are amoeboid protists which produce intricate mineral skeletons. The skeletons, usually of silica (SiO2), have a central capsule. this divides the cell into inner and outer portions, called endoplasm and ectoplasm.

Radiolaria are found as zooplankton throughout the ocean, and their skeletal remains cover large portions of the ocean bottom as radiolarian ooze. Radiolarians have existed since the beginning of the Paleozoic era.[1] They take their name from the radial symmetry characteristic of many species. They are valuable markers or diagnostic fossils, because they have a rapid turn-over of species. This helps to identify strata, and to correlate strata at different sites.

Radiolaria are heterotrophs (eat other things), but also may include protist algae as endosymbionts.

Structure

The skeletons of radiolarians are generally organized around spicules, or spines, which extend from the main skeletal mass. Formed from the fusion of many of these spines is the outermost skeleton, the shell or test. Connecting this shell to the many concentrically organized inner shells are bars or beams, which strengthen and support the structure.[2]

The nuclei and most other organelles are in the endoplasm, while the ectoplasm is filled with frothy vacuoles and lipid droplets, keeping them buoyant. Often it also contains symbiotic algae which provide much of the cell's energy. Not much is known about the living amoebae because they cannot be cultured in the laboratory.

Other links

  • NHM rotating glass model [1]]

References


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